Note

Taxa Associated with the Family Sepiidae Keferstein, 1866

(data as of June 2002)

Nominal Genus-Level Taxa

Acanthosepion Rochebrune, 1884b:100.

CURRENT SYSTEMATIC STATUS. -- Species complex of Sepia [fide Khromov et al. (1998:80)]
TYPE SPECIES. -- Sepia aculeata Van Hasselt, 1835 In Ferussac and D'Orbigny, 1834-1848 by subsequent designation of Adam and Rees (1966:147)

Amplisepia Iredale, 1926a:194.

CURRENT SYSTEMATIC STATUS. -- Sepia Linne, 1758 [fide Adam and Rees (1966:34)]
TYPE SPECIES. -- Sepia apama Gray, 1849 by original designation

Andreasepia Grimpe, 1922:44.

CURRENT SYSTEMATIC STATUS. -- Sepia Linne, 1758 [fide Nesis (1987a:94)]
TYPE SPECIES. -- Sepia andreana Steenstrup, 1875 by original designation

(Anomalosepia) Khromov, 1987b:1172.

CURRENT SYSTEMATIC STATUS. -- Species complex of Sepia [fide Khromov et al. (1998:81)]
TYPE SPECIES. -- Sepia omani Adam and Rees, 1966 by monotypy

Arctosepia Iredale, 1926a:193.

CURRENT SYSTEMATIC STATUS. -- Generic affinity undetermined [fide Adam and Rees (1966:81)]
TYPE SPECIES. -- Arctosepia limata Iredale, 1926 by original designation

Ascarosepion Rochebrune, 1884b:98.

CURRENT SYSTEMATIC STATUS. -- Sepia Linne, 1758 [fide Adam and Rees (1966:47)]
TYPE SPECIES. -- Ascarosepion verreauxi Rochebrune, 1884 by subsequent designation of Hoyle (1910b:408)

Aurosepina Jothinayagam, 1987:32.

CURRENT SYSTEMATIC STATUS. -- Unresolved
TYPE SPECIES. -- Sepia arabica Massy, 1916 by monotypy

Blandosepia Iredale, 1940:442.

CURRENT SYSTEMATIC STATUS. -- Sepia Linne, 1758 [fide Adam and Rees (1960:105)]
TYPE SPECIES. -- Blandosepia baxteri Iredale, 1940 by monotypy

Chondrosepia Ruppell and Leuckart, 1828:21.

CURRENT SYSTEMATIC STATUS. -- Sepioteuthis Blainville, 1824 [fide Ehrenberg (1831:[3])]
TYPE SPECIES. -- Chondrosepia loliginiformis Ruppell and Leuckart, 1828 by monotypy

Crumenasepia Iredale, 1926b:239.

CURRENT SYSTEMATIC STATUS. -- Sepia Linne, 1758 [fide Adam and Rees (1966:22)]
TYPE SPECIES. -- Crumenasepia hulliana Iredale, 1926 by original designation

Decorisepia Iredale, 1926a:193.

CURRENT SYSTEMATIC STATUS. -- Sepia Linne, 1758 [fide Lu (1998a:180)]
TYPE SPECIES. -- Decorisepia rex Iredale, 1926 by original designation

Diphtherosepion Rochebrune, 1884b:79.

CURRENT SYSTEMATIC STATUS. -- Sepiella Gray, 1849 [fide Adam and Rees (1966:148)]
TYPE SPECIES. -- Sepia ornata Rang, 1837 by subsequent designation of Hoyle (1910b:411)

Doratosepion Rochebrune, 1884b:95.

CURRENT SYSTEMATIC STATUS. -- Species complex of Sepia [fide Khromov et al. (1998:81)]
TYPE SPECIES. -- Sepia andreana Steenstrup, 1875 by subsequent designation of Hoyle (1910b:409)

(Eusepia) Naef, 1923:545.

CURRENT SYSTEMATIC STATUS. -- Sepia Linne, 1758 [fide Adam and Rees (1966:148)]
TYPE SPECIES. -- Sepia officinalis Linne, 1758 by original designation

(Fiscisepia) Iredale, 1926b:239.

CURRENT SYSTEMATIC STATUS. -- Sepia Linne, 1758 [fide Adam and Reese (1966:14)]
TYPE SPECIES. -- Acanthosepion ellipticum adjacens Iredale, 1926 by original designation

Glyptosepia Iredale, 1926a:191.

CURRENT SYSTEMATIC STATUS. -- Sepia Linne, 1758 [fide Lu (1998a:175)]
TYPE SPECIES. -- Glyptosepia opipara Iredale, 1926 by original designation

Hemisepion Fischer, 1882 In 1880-1887:357.

Emendation of Hemisepius Steenstrup, 1875

Hemisepion Rochebrune, 1884b:78.

"Emendation [?] of Hemisepius" [fide Hoyle (1910b:409)]

Hemisepius Steenstrup, 1875:471.

CURRENT SYSTEMATIC STATUS. -- Species complex of Sepia [fide Khromov et al. (1998:81)]
TYPE SPECIES. -- Hemisepius typicus Steenstrup, 1875 by monotypy

Lophosepion Rochebrune, 1884b:90.

CURRENT SYSTEMATIC STATUS. -- Sepia Linne, 1758 [fide Adam and Rees (1966:100)]
TYPE SPECIES. -- Sepia lefebrei D'Orbigny, 1839-1842 In Ferussac and D'Orbigny, 1834-1848 by subsequent designation of Adam and Rees (1966:148)]

Mesembrisepia Iredale, 1926a:190.

CURRENT SYSTEMATIC STATUS. -- Sepia Linne, 1758 [fide Lu (1998a:174)]
TYPE SPECIES. -- Mesembrisepia macandrewi Iredale, 1926 by original designation

(Metasepia) Hoyle, 1885c:199.

CURRENT SYSTEMATIC STATUS. -- Valid genus [fide Lu (1998a:186)]
TYPE SPECIES. -- Sepia (Metasepia) pfefferi Hoyle, 1885 by monotypy

(Parasepia) Naef, 1923:545.

CURRENT SYSTEMATIC STATUS. -- Sepia Linne, 1758 [fide Nesis (1987a:94)]
TYPE SPECIES. -- Sepia orbignyana Ferussac In D'Orbigny, 1826 by original designation

(Platysepia) Naef, 1923:546.

CURRENT SYSTEMATIC STATUS. -- Sepia Linne, 1758 [fide Nesis (1987a:94)]
TYPE SPECIES. -- Sepia esculenta Hoyle, 1885 by orginal designation

Ponderisepia Iredale, 1926b:239.

CURRENT SYSTEMATIC STATUS. -- Sepia Linne, 1758 [fide Adam and Rees (1966:33)]
TYPE SPECIES. -- Ponderisepia eclogaria Iredale, 1926 by original designation

Rhombosepion Rochebrune, 1884b:82.

CURRENT SYSTEMATIC STATUS. -- Species complex of Sepia [fide Khromov et al. (1998:80)]
TYPE SPECIES. -- Sepia rupellaria D'Orbigny, 1834 In Ferussac and D'Orbigny, 1834-1848 by subsequent designation of Hoyle (1910b:412)

Sepia LinnÈ, 1758:658.

CURRENT SYSTEMATIC STATUS. -- Valid genus [fide Adam and Rees (1966)]
TYPE SPECIES. -- Sepia officinalis Linne, 1758 by elimination [fide Hoyle (1910b:412)]

Sepia Schneider, 1784:109.

CURRENT SYSTEMATIC STATUS. -- "never published"; rejected generic name [fide Opinion 233, ICZN (1954:277)]

(Sepiella) Gray, 1849:106.

CURRENT SYSTEMATIC STATUS. -- Valid genus [fide Khromov et al. (1998:133)]
TYPE SPECIES. -- Sepia ornata Rang, 1837 by subsequent designation of Hoyle (1910b:412)]

Solitosepia Iredale, 1926a:188.

CURRENT SYSTEMATIC STATUS. -- Sepia Linne, 1758 [fide Lu (1998a:171)]
TYPE SPECIES. -- Solitosepia liliana Iredale, 1926 by original designation

Spathidosepion Rochebrune, 1884b:93.

CURRENT SYSTEMATIC STATUS. -- Sepia Linne, 1758 [fide Adam and Rees (1966:106)]
TYPE SPECIES. -- Sepia tuberculata Lamarck, 1798 by subsequent designation of Hoyle (1910b:413)]

Tenuisepia Cotton, 1932:546.

CURRENT SYSTEMATIC STATUS. -- Sepia Linne, 1758 [fide Adam and Rees (1966:87)]
TYPE SPECIES. -- Tenuisepia mira Cotton, 1932 by original designation

Nominal Species-Level Taxa (As Introduced Binomial)

Sepia aculeata Van Hasselt, 1835
In Ferussac and D'Orbigny, 1834-1848:287, Seiche pls 5bis, 25
[fide Tillier and Boucher-Rodoni (1994:101); taxon dated from pl 5bis]

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:82)]
TYPE REPOSITORY. -- MNHN Syntype 1-6-186 and 1-6-187 (sepion) [fide Lu et al. (1995:310)]
TYPE LOCALITY. -- Java

Sepia acuminata Smith, 1916:21, pl 2 figs 3-4.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:82)]
TYPE REPOSITORY. -- BMNH Syntypes (2) 1890.9.22.332-333 (sepion) [fide Lipinski et al. (2000:102)]
TYPE LOCALITY. -- Port Elizabeth; Tongaat Beach, Natal, South Africa [fide Adam and Rees (1966:53)]

Sepia adami Roeleveld, 1972:224, fig 10.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:83)]
TYPE REPOSITORY. -- SAM Holotype A-31394 [fide Khromov et al. (1998:83)
TYPE LOCALITY. -- S 79∞ E off Cape Natal, South Africa

Solitosepia plangon adhaesa Iredale, 1926b:238.

CURRENT SYSTEMATIC STATUS. -- Sepia plangon Gray, 1849 [fide Lu (1998a:178)]
TYPE REPOSITORY. -- AMS Holotype C133304 [fide Rudman (1983:68)]
TYPE LOCALITY. -- North-west Island, Queensland, Australia

Acanthosepion (Fiscisepia) ellipticum adjacens Iredale, 1926b:239, pl 35 figs 5-6.

CURRENT SYSTEMATIC STATUS. -- Sepia elliptica Hoyle, 1885 [fide Adam and Reese (1966:14)]
TYPE REPOSITORY. -- AMS Holotype C133302 [fide Rudman (1983:68)]
TYPE LOCALITY. -- North-west Island, Queensland, Australia

Sepia affinis D'Orbigny, 1826:156.

CURRENT SYSTEMATIC STATUS. --?Sepioteuthis sepioidea (Blainville, 1823) (pars) [fide Adam and Rees (1966:149)]
TYPE REPOSITORY. -- MNHN Type; specimen not extant [fide Lu et al. (1995:310)]
TYPE LOCALITY. -- “Le cap de Bonne-Esperance, Monfort”

Sepia affinis Souleyet, 1852:35, pl 3 figs 13-14.

CURRENT SYSTEMATIC STATUS. -- Sepiella inermis Van Hasselt, 1835 In Ferussac and D'Orbigny, 1834-1848 [fide Adam and Rees (1966:124)]
TYPE REPOSITORY. -- MNHN Syntype 1-7-283 [fide Lu et al. (1995:310)]
TYPE LOCALITY. -- Bay of Tourane [Da Nang, Vietnam]

Sepia kobiensis albatrossi Sasaki, 1920:195, pl 26 figs 2-3.

CURRENT SYSTEMATIC STATUS. -- Sepia kobiensis Hoyle, 1885 [fide Adam and Rees (1966:71)]
TYPE REPOSITORY. -- NMNH Holotype 332862a [fide Roper and Sweeney (1978:3)]
TYPE LOCALITY. -- Okinoshima, Krusenstern Strait (34∞19'N, 130∞09'E, Sea of Japan) [fide Sasaki (1929:216)]

Sepia andreana Steenstrup, 1875:474, pl 1 figs 11-19.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:84)]
TYPE REPOSITORY. -- ZMUC Syntypes [fide Kristensen and Knudsen (1983:220)]; MNHN Lectotype [fide Rochebrune (1884b:96) and Lu et al. (1995:310)]
TYPE LOCALITY. -- Hakodate, Japan

Sepia andreanoides Hoyle, 1885c:193.

CURRENT SYSTEMATIC STATUS. -- Sepia kobiensis Hoyle, 1885 [fide Khromov et al. (1998:101)]
TYPE REPOSITORY. -- BMNH Syntypes (2) 1889.4.24.78-79 [fide Lipinski et al. (2000:102)]; NMGW Syntype (1)
TYPE LOCALITY. -- Yokohama market, Japan [fide Adam and Rees (1966:71)]

Sepia angulata Roeleveld, 1972:242, pls 44d, 45a-d.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:84)]
TYPE REPOSITORY. -- SAM Holotype A-31317 (sepion only)
TYPE LOCALITY. -- Bloubergstrand [33∞48'S, 18∞27'E], South Africa

Sepia antillarum D'Orbigny, 1841 In 1841-1853:33.

CURRENT SYSTEMATIC STATUS. -- Doubtful species [fide Adam and Rees (1966:149)]
TYPE REPOSITORY. -- MNHN Syntypes (2); specimens not extant [fide Lu et al. (1995:310)]
TYPE LOCALITY. -- Jamaica and Martinique

Sepia apama Gray, 1849:103.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Lu (1998a:160)]
TYPE REPOSITORY. -- BMNH Holotype 1846.5.25.2 (sepion) [fide Lipinski et al. (2000:102)]
TYPE LOCALITY. -- Port Adelaide [34∞50'S, 138∞30'E], Australia

Sepia appellofi Wulker, 1910:14, figs 8, 15-18.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:84)]
TYPE REPOSITORY. -- ZSM Holotype [fide Khromov et al. (1998:84)]
TYPE LOCALITY. -- Misaki, Japan

Sepia arabica Massy, 1916:228, pl 23-24.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:85)]
TYPE REPOSITORY. -- ZSI Syntypes M8113/1, M8122/1
TYPE LOCALITY. -- Lacadive Sea and Persian Gulf

Sepia (Doratosepia) aureomaculata Okutani and Horikawa, 1987:62, 1 fig.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:86)]
TYPE REPOSITORY. -- NSMT Holotype Mo-64224 [fide Kubodera and Tsuchiya (1993:unpaginated)]
TYPE LOCALITY. -- Tosa Bay, Japan

Sepia australis Quoy and Gaimard, 1832:70, pl 5 figs 3-7.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:87)]
TYPE REPOSITORY. -- MNHN Syntype 1-6-211 [fide Lu et al. (1995:311)]
TYPE LOCALITY. -- Banc des Aiguilles, Cape of Good Hope

Sepia australis D'Orbigny, 1835
In Ferussac and D'Orbigny, 1834-1848:285, Seiche pl 7 fig 4
[fide Tillier and Boucher-Rodoni (1994:101); taxon dated from plate]

CURRENT SYSTEMATIC STATUS. -- Junior homonym of Sepia australis Quoy and Gaimard, 1832
TYPE REPOSITORY. -- MNHN Syntype 1-6-194 (sepion) [fide Lu et al. (1995:311)]
TYPE LOCALITY. -- Grand Ocean, New Holland [Australia], Kangaroo Island [fide Lu et al. (1995:311)]

Sepia bandensis Adam, 1939b:1, figs 1-4.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:87)]
TYPE REPOSITORY. -- IRSNB Syntypes IG-10542 and 10652 [fide Khromov et al. (1998:87)]
TYPE LOCALITY. -- Banda Neira (Indonesia)

Blandosepia bartletti Iredale, 1954:67, pl 5 figs 15-16.

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepia bartletti [fide Khromov et al. (1998:87)]
TYPE REPOSITORY. -- AMS Holotype C133318 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Louisiade Archipelago

Sepia (Doratosepion) bathyalis Khromov, Nikitina and Nesis, 1991:18, pl 3.

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepia bathyalis [fide Khromov et al. (1998:87)]
TYPE REPOSITORY. -- ZMMGU Holotype Y-361 [fide Ivanov and Sysoev (2000:108)]
TYPE LOCALITY. -- SW Madagascar

Blandosepia baxteri Iredale, 1940:442.

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepia baxteri [fide Adam and Rees (1966:105)]
TYPE REPOSITORY. -- AMS Holotype C133317 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Lord Howe Island

Sepia kobiensis beppuana Sasaki, 1929:211, text-fig 116, pl 19 figs 13-15.

CURRENT SYSTEMATIC STATUS. -- Sepia kobiensis Hoyle, 1885 [fide Adam and Rees (1966:71)]
TYPE REPOSITORY. -- MSUT[TIU] Syntypes
TYPE LOCALITY. -- Beppu, Bungo Province, Japan

Sepia bertheloti D'Orbigny, 1835
In Ferussac and D'Orbigny, 1834-1848:274, Seiche pls 11, 23
[fide Tillier and Boucher-Rodoni (1994:101); taxon dated from plate 11]

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:88)]
TYPE REPOSITORY. -- MNHN Syntypes 1-4-75 (without sepion), 1-4-76 (without sepion), 1-4-77 (3 sepions) [fide Lu et al. (1995:312)]; IRSNB Syntype [fide F.G. Hochberg (pers. comm.)]; BMNH Syntype (sepion) 1854.9.28.1 [fide Lipinski et al. (2000:102)]
TYPE LOCALITY. -- Teneriffe, Canary Islands

Sepia bidhaia Reid, 2000:4, figs 2-7.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide; taxon not yet reviewed]
TYPE REPOSITORY. -- NMV Holotype F82478
TYPE LOCALITY. -- Queensland, Australia

Sepia bilineata Quoy and Gaimard, 1832:66, pl 2 fig 1.

CURRENT SYSTEMATIC STATUS. -- Species dubium [fide Lu and Tait (1983:199)]
TYPE REPOSITORY. -- MNHN Type; specimen not extant† [fide Lu et al. (1995:312)]
TYPE LOCALITY. -- New Holland [Australia], Port Western

Sepia biserialis Blainville, 1827:284.

CURRENT SYSTEMATIC STATUS. -- Sepia elegans Blainville, 1827 [fide Adam and Rees (1966:94)]
TYPE REPOSITORY. -- Unresolved; [other Blainville types at MNHN] [not reported by Lu et al. (1995)]
TYPE LOCALITY. -- Mediterranean

Sepia bisserialis Verany, 1851:73, pl 26.

CURRENT SYSTEMATIC STATUS. -- Sepia elegans Blainville, 1827 [fide Adam and Rees (1966:94)]
TYPE REPOSITORY. -- Unresolved; MHNN? Syntypes [other Verany taxa there fide G.L. Voss (pers. comm.)]
TYPE LOCALITY. -- Mediterranean Sea; Adriatic Sea

Sepia blainvillei D'Orbigny, 1839-1842
In Ferussac and D'Orbigny, 1834-1848:288, Seiche pl 21
[fide Tillier and Boucher-Rodoni (1994:101); taxon dated from plate]

CURRENT SYSTEMATIC STATUS. -- Sepia aculeata Van Hasselt, 1835 In Ferussac and D'Orbigny, 1834-1848 [fide Adam and Rees (1966:12)]
TYPE REPOSITORY. -- MNHN Type; specimen not extant [fide Lu et al. (1995:312)]
TYPE LOCALITY. -- Bombay, Grand Ocean [India] [fide Lu et al. (1995:312)]

Sepia boscii Lesueur, 1821:101.

Nomen nudum; [see Lu et al. (1995:312)]

Sepia brachycheira Tapparone-Canefri, 1877:278.

CURRENT SYSTEMATIC STATUS. -- Doubtful species [fide Adam and Rees (1966:149)]
TYPE REPOSITORY. -- MSNG Holotype
TYPE LOCALITY. -- Northwest coast of Papua New Guinea

Sepia braggi Verco, 1907:213, pl 27 fig 6.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Reid (2000:13)]
TYPE REPOSITORY. -- SAMA Holotype D14130 (sepion) [fide Zeidler and MacPhail (1978:382)]
TYPE LOCALITY. -- Glenelg, South Australia [fide Lu (1998a:161)]

Sepia brevimana Steenstrup, 1875:475.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:89)]
TYPE REPOSITORY. -- ZMUC Lectotype [fide Kristensen and Knudsen (1983:221)]
TYPE LOCALITY. -- Indian Ocean

Sepia burnupi Hoyle, 1904a:27, pl 1.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:89)]
TYPE REPOSITORY. -- BMNH Holotype 1905.1.19.5 (sepion) [fide Lipinski et al. (2000:102)]
TYPE LOCALITY. -- Natal, South Africa

Sepia capensis D'Orbigny, 1835
In Ferussac and D'Orbigny, 1834-1848:278, Seiche pls 7, 12, 17
[fide Tillier and Boucher-Rodoni (1994:101); taxon dated from pl 7]

CURRENT SYSTEMATIC STATUS. -- Sepia australis Quoy and Gaimard, 1832 [fide Voss (1962d:246)]
TYPE REPOSITORY. -- MNHN Type; specimen not extant [fide Lu et al. (1995:313)]
TYPE LOCALITY. -- Banc des Aiguilles, Cape of Good Hope

Sepia carinata Sasaki, 1920:192, pl 25 fig 2, pl 26 fig 1.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:90)]
TYPE REPOSITORY. -- NMNH Holotype 332849 [fide Roper and Sweeney (1978:5)]
TYPE LOCALITY. -- Sagami Sea, Japan (35∞04'50"N, 139∞38'18"E)

Sepia chirotrema Berry, 1918:268, text-figs 56-61, pls 74-79.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Lu (1998a:164)]
TYPE REPOSITORY. -- AMS Holotype; specimen not extant [fide Rudman (1983:67)]
TYPE LOCALITY. -- Investigator Strait, Australia [fide Sweeney et al. (1988:12)]

Sepia chrysophtalmos Tilesius, 1813:pl 38 figs 32-33.

CURRENT SYSTEMATIC STATUS. --?Idiosepius paradoxus (Ortmann, 1888) [fide Berry (1932:52)]
TYPE REPOSITORY. -- Unresolved [see also Tryon (1879:164, pl 69 fig 256)]
TYPE LOCALITY. -- Japan (on Fucus) [fide D’Orbigny (1845:373)]

Hemisepius typicus chuni Thore, 1945:52.

CURRENT SYSTEMATIC STATUS. -- Sepia (Hemisepius) typicus (Steenstrup, 1875) [fide Adam and Rees (1966:117)]
TYPE REPOSITORY. -- Unresolved; ZMB? Holotype [species not traced in Glaubrecht and Salcedo-Vargas (2000)]
TYPE LOCALITY. -- St. Francis Bay, Agulhas Bank, 34∞9'S, 24∞59'E [fide Chun (1915:412)]

Sepia confusa Smith, 1916:24, pl 2 figs 7-8.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:90)]
TYPE REPOSITORY. -- BMNH Holotype 1920.3.23.1 (sepion) [fide Lipinski et al. (2000:103)]
TYPE LOCALITY. -- Tongaat Beach, Natal [fide Adam and Rees (1966:65)]

Decorisepia cottesloensis Cotton, 1929:90, pl 16.

CURRENT SYSTEMATIC STATUS. -- Sepia rex (Iredale, 1926) [fide Adam (1979)]
TYPE REPOSITORY. -- SAMA Holotype D13681 [fide Zeidler and MacPhail (1978:381)]
TYPE LOCALITY. -- Cottesloe, Western Australia

Sepia cottoni Adam, 1979:193, pl 11 figs 1-6.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Reid (2000:21)]
TYPE REPOSITORY. -- WAM Holotype 435.65 [fide Slack-Smith (1983:74)]
TYPE LOCALITY. -- West of Lancelin Island, Western Australia [fide Lu (1998a:164)]

Sepia kobiensis crassa Sasaki, 1929:213, pl 19 figs 16-18.

CURRENT SYSTEMATIC STATUS. -- Sepia kobiensis Hoyle, 1885 [fide Adam and Rees (1966:71)]
TYPE REPOSITORY. -- MSUT[TIU] Syntypes (7)
TYPE LOCALITY. -- Enoura, Suruga Province, Japan

Sepia cultrata Hoyle, 1885c:198.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Lu (1998a:166)]
TYPE REPOSITORY. -- BMNH Holotype 1889.4.24.75 [fide Lipinski et al. (2000:103)]
TYPE LOCALITY. -- 36∞59'S, 150∞20'E, off Twofold Bay, SE Australia

Sepia cyanomarginata Ruppell In Wulker, 1920:54, 55.

Nomen nudum.

Diphtherosepion dabryi Rochebrune, 1884b:81.

CURRENT SYSTEMATIC STATUS. -- Sepia esculenta Hoyle, 1885 and Sepiella japonica Sasaki, 1929 [fide Adam (1944:221)]
TYPE REPOSITORY. -- MNHN Syntypes (2) 1-6-220, 1-7-260 and 1-7-261 (sepion) [fide Lu et al. (1995:314)]
TYPE LOCALITY. -- Canton, China

Sepia dannevigi Berry, 1918:264, text-figs 51-55, pls 73-74.

CURRENT SYSTEMATIC STATUS. -- Sepia hedleyi Berry, 1918 [fide Reid (2001b:83)]
TYPE REPOSITORY. -- AMS Holotype C148249 [fide Sweeney et al. (1988:13)]
TYPE LOCALITY. -- South of Kangaroo Island, South Australia [fide Lu (1998a:168)]

Sepia dollfusi Adam, 1941d:12, pl 2 fig 3.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:92)]
TYPE REPOSITORY. -- MNHN Holotype 1-6-182 [fide Lu et al. (1995:315)]
TYPE LOCALITY. -- Perim, Red Sea

Sepia (Hemisepius) dubia Adam and Rees, 1966:119, text-fig 1, pls 34, 36.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:93)]
TYPE REPOSITORY. -- BMNH Holotype 1963.103W [fide Lipinski et al. (2000:103)]
TYPE LOCALITY. -- False Bay, South Africa

Ponderisepia eclogaria Iredale, 1926b:239, pl 35 figs 7-8.

CURRENT SYSTEMATIC STATUS. -- Sepia latimanus Quoy and Gaimard, 1832 [fide Adam and Rees (1966:33)]
TYPE REPOSITORY. -- AMS Holotype C19085 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Masthead Island, Queensland, Australia

Sepia elegans Blainville, 1827:284.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:93)]
TYPE REPOSITORY. -- MNHN Syntypes; specimens not extant [fide Lu et al. (1995:315)]
TYPE LOCALITY. -- Sicily, Italy

Sepia elliptica Hoyle, 1885c:189.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Lu (1998a:168)]
TYPE REPOSITORY. -- Eight syntypes: BMNH Syntypes (4) 1889.4.24.71-72 and 1889.4.24.73-74 [fide Lipinski et al. (2000:103)]; NMGW Syntypes [?]
TYPE LOCALITY. -- 9∞59'S, 139∞42'E and 8∞56'S, 136∞05'E (Arafura Sea)

Sepia elobyana Adam, 1941b:121, text-fig 12, pl 13.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:94)]
TYPE REPOSITORY. -- ZMH Holotype E.K.7453 [fide Khromov et al. (1998:94)]; no longer extant [fide Guerrero-Kommritz (2001:174)]
TYPE LOCALITY. -- Eloby Island, Gulf of Guinea [fide Adam and Rees (1966:103)]

Sepia elongata D'Orbigny, 1839-1842
In Ferussac and D'Orbigny, 1834-1848:283, Seiche pl 24 figs 7-10
[fide Tillier and Boucher-Rodoni (1994:101); taxon dated from plate]

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:94)]
TYPE REPOSITORY. -- MNHN Lectotype 1-6-197 (sepion) [fide Rochebrune (1884b:98)]
TYPE LOCALITY. -- Cosseir, Red Sea [fide Lu et al. (1995:315)]

Acanthosepion enoplon Rochebrune, 1884b:108.

CURRENT SYSTEMATIC STATUS. -- Sepia orbignyana Ferussac In D'Orbigny, 1826 [fide Adam and Rees (1966:150)]
TYPE REPOSITORY. -- MNHN Holotype 1-4-85 [fide Lu et al. (1995:315)]
TYPE LOCALITY. -- Naples, Italy

Sepia erostrata Sasaki, 1929:183, pl 17 figs 7-9.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:95)]
TYPE REPOSITORY. -- MSUT[TIU] Holotype
TYPE LOCALITY. -- Manzuru, Sagami Province, Japan

Sepia esculenta Hoyle, 1885c:188.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:95)]
TYPE REPOSITORY. -- BMNH Syntypes 1889.4.24.69-70 [fide Lipinski et al. (2000:103)]
TYPE LOCALITY. -- Purchased at market, Yokohama, Japan

Sepia exsignata Barnard, 1962:250, fig 3.

CURRENT SYSTEMATIC STATUS. -- Sepia burnupi Hoyle, 1904 [fide Khromov et al. (1998:89)]
TYPE REPOSITORY. -- SAM Holotype A6525
TYPE LOCALITY. -- Off Umhlanga River, Natal [South Africa]

Sepia faurei Roeleveld, 1972:251, figs 15-16.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:97)]
TYPE REPOSITORY. -- SAM Holotype A-30144
TYPE LOCALITY. -- S 14o E of Cape Seal [34∞07'S, 23∞24'E], South Africa

Sepia filliouxi Lafont, 1869:11.

CURRENT SYSTEMATIC STATUS. -- Sepia officinalis Linne, 1758 [fide Khromov et al. (1998:81)]
TYPE REPOSITORY. -- Unresolved; Museum de Bordeaux Syntypes (2) [fide Lagatu (1888:112-114)
TYPE LOCALITY. -- Gulf of Gascogne

Sepia fillouxii Lafont, 1868:121.

Nomen nudum.

Sepia fischeri Lafont, 1871:271.

CURRENT SYSTEMATIC STATUS. --?Sepia filliouxi Lafont, 1869 [fide Adam and Rees (1966:150)] = Sepia officinalis Linne, 1758 [fide Khromov et al. (1998:81)]
TYPE REPOSITORY. -- Syntypes (2): male, Musee de la Societe scientifique d’Arachon; female, Museum de Bordeaux [fide Lagatu (1888:109-110)]
TYPE LOCALITY. -- Bay of Biscay, France

Sepia (Doratosepion) foliopeza Okutani and Tagawa, 1987:67, 5 figs.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:97)]
TYPE REPOSITORY. -- NSMT Holotype Mo-64232 [fide Kubodera and Tsuchiya (1993:unpaginated)]
TYPE LOCALITY. -- East China Sea

Sepia formosana Berry, 1912b:420, text-fig 2, pl 9 fig 7.

CURRENT SYSTEMATIC STATUS. -- Sepia pharaonis Ehrenberg, 1831 [fide Adam and Rees (1966:22)]
TYPE REPOSITORY. -- CASIZ Holotype 021668 [fide Sweeney et al. (1988:13)]
TYPE LOCALITY. -- Takao fish market, Formosa [Taiwan]

Ascarosepion singhalensis foxi Robson, 1927b:325, figs 79-81.

CURRENT SYSTEMATIC STATUS. -- Sepia pharaonis Ehrenberg, 1831 [fide Adam and Rees (1966:22)]
TYPE REPOSITORY. -- BMNH Holotype 1926.4.8.6 [fide Lipinski et al. (2000:105)]
TYPE LOCALITY. -- Kabret, Suez Canal

Sepia framea Ortmann, 1891:675, pl 46 fig 2a-d.

CURRENT SYSTEMATIC STATUS. -- Sepia pharaonis Ehrenberg, 1831 [fide Adam and Rees (1966:22)]
TYPE REPOSITORY. -- MZUS Holotype [fide Johnson (1977:237)]
TYPE LOCALITY. -- Ceylon [Sri Lanka]

Sepia galei Meyer, 1909:332, figs 4-6.

CURRENT SYSTEMATIC STATUS. --?Sepia papuenis Hoyle, 1885 [fide Lu (1998a:175)]
TYPE REPOSITORY. -- ZMH Holotype (sepion missing) 3841 [fide Guerrero-Kommritz (2001:171)]
TYPE LOCALITY. -- Shark Bay, near Brown Station on Dirk Hartog [Australia] [fide Adam (1979:169)]

Glyptosepia gemellus Iredale, 1926a:192, pl 22.

CURRENT SYSTEMATIC STATUS. -- Sepia cultrata Hoyle, 1885 [fide Lu (1998a:166)]
TYPE REPOSITORY. -- AMS Holotype C133306 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Manly Beach, N.S.W., Australia

Solitosepia genista Iredale, 1954:66, pl 5 figs 17-18.

CURRENT SYSTEMATIC STATUS. -- Sepia papuensis Hoyle, 1885 [fide Lu (1998a:175)]
TYPE REPOSITORY. -- AMS Holotype C133309 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Broome, Western Australia

Sepia gibba Ehrenberg, 1831:[5].

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:98)]
TYPE REPOSITORY. -- ZMB Holotype 92 [fide Khromov et al. (1998:98)] [species not traced in Glaubrecht and Salcedo-Vargas (2000)]
TYPE LOCALITY. -- Red Sea, near Hama [fide Adam and Rees (1966:100)]

Sepia gibbosa D'Orbigny, 1845 In 1845-1847:287.

Incorrect subsequent spelling of Sepia gibba Ehrenberg, 1831.

Sepia gigas Oken, 1815 In 1813-1826:344.

CURRENT SYSTEMATIC STATUS. -- "Poulpe colossal" [see Adam and Rees (1966:151)]
TYPE REPOSITORY. -- Unresolved
TYPE LOCALITY. -- Unresolved

Solitosepia glauerti Cotton, 1929:87, pl 14.

CURRENT SYSTEMATIC STATUS. -- Sepia chirotrema Berry, 1918 [fide Lu (1998a:164)]
TYPE REPOSITORY. -- SAMA Holotype D13628 (sepion) [fide Zeidler and MacPhail (1978:382)]
TYPE LOCALITY. -- Rottnest Island, Western Australia

Acanthosepion goreense Rochebrune, 1884b:109.

CURRENT SYSTEMATIC STATUS. -- Sepia hierredda Rang, 1835 In Ferussac and D'Orbigny, 1834-1848 [fide Khromov et al. (1998:99)]
TYPE REPOSITORY. -- MNHN Holotype 1-3-35 and 1989 (sepion) [fide Lu et al. (1995:316)]
TYPE LOCALITY. -- Goree, Cotes de Senegambie, Senegal [fide Lu et al. (1995)]

Sepia grahami Reid, 2001a:160, figs 1-5.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide; taxon not yet reviewed]
TYPE REPOSITORY. -- NMV Holotype F87757
TYPE LOCALITY. -- New South Wales, Australia

Sepia groenlandica Dewhurst, 1834:263.

CURRENT SYSTEMATIC STATUS. -- Nomen nudum; ?Bathypolypus arcticus (Prosch, 1847) [fide Adam and Rees (1966:151)]
TYPE REPOSITORY. -- No type designated
TYPE LOCALITY. -- Greenland Seas

Sepia harmeri Robson, 1928c:8, figs 1d, 2-3.

CURRENT SYSTEMATIC STATUS. -- Sepia latimanus Quoy and Gaimard, 1832 [fide Adam and Rees (1966:33)]
TYPE REPOSITORY. -- Unresolved; MNHN? Holotype [not trace by Lu et al. (1995)]
TYPE LOCALITY. -- Poulo-Condor, Indo-China

Acanthosepion hasselti Rochebrune, 1884b:101.

CURRENT SYSTEMATIC STATUS. -- Sepia aculeata Van Hasselt, 1835 In Ferussac and D'Orbigny, 1834-1848† [fide Adam and Rees (1966:12)]
TYPE REPOSITORY. -- MNHN Type; specimen not extant [fide Lu et al. (1995:317)]
TYPE LOCALITY. -- Indian Sea

Sepia hedleyi Berry, 1918:258, text-figs 48-50, pls 71-72.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Reid (2001b:83)]
TYPE REPOSITORY. -- AMS Holotype C148252 [fide Sweeney et al. (1988:14)]
TYPE LOCALITY. -- Investigator Strait, Australia [fide Lu (1998a:169)]

Solitosepia hendryae Cotton, 1929:87, pl 15.

CURRENT SYSTEMATIC STATUS. -- Sepia chirotrema Berry, 1918 [fide Lu (1998a:164)]
TYPE REPOSITORY. -- SAMA Holotype D13625 (sepion) [fide Zeidler and MacPhail (1978:382)]
TYPE LOCALITY. -- Rottnest Island, Western Australia

Sepia hercules Pilsbry, 1894:144.

CURRENT SYSTEMATIC STATUS. -- Sepia latimanus Quoy and Gaimard, 1832 [fide Adam and Rees (1966:33)]
TYPE REPOSITORY. -- ANSP Syntype 66090 [fide Spamer and Bogan (1992:2)]
TYPE LOCALITY. -- Loo Choo Islands (Ryukyu Islands)

Sepia hexapodia Molina, 1782:199, 348.

CURRENT SYSTEMATIC STATUS. -- "insecte" [fide D'Orbigny (1845:300 In 1845-1847); see Adam and Rees (1966:151)]

Sepiella heylei Sasaki, 1929:text-fig 170.

Nomen nudum; illustration for Sepiella japonica Sasaki, 1929 [fide Adam and Rees (1966:128)]

Rhombosepion hieronis Robson, 1924c:645, pl 2 figs 9-11.

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepia hieronis [fide Adam and Rees (1966:112)]
TYPE REPOSITORY. -- BMNH Syntypes (4) 1924.9.8.54-56 [fide Lipinski et al. (2000:103)]
TYPE LOCALITY. -- Off Cape Town, South Africa [fide Khromov et al. (1998:98)]

Sepia hierredda Rang, 1835
In Ferussac and D'Orbigny, 1834-1848:268, pls. 13, 18
[fide Tillier and Boucher-Rodoni (1994:101); taxon dated from plate 13]

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:99)]
TYPE REPOSITORY. -- MNHN Syntypes 1-4-66 and 1-6-195 [fide Lu et al. (1995:317)]
TYPE LOCALITY. -- Goree, Senegal

Sepia hoylei Ortmann, 1888:650, pls 22-23.

CURRENT SYSTEMATIC STATUS. -- Sepia esculenta Hoyle, 1885 [fide Adam and Rees (1966:18)]
TYPE REPOSITORY. -- MZUS Syntypes [fide Johnson (1977:237)]
TYPE LOCALITY. -- Japan, several locations

Crumenasepia hulliana Iredale, 1926b:239, pl 35 figs 1-2.

CURRENT SYSTEMATIC STATUS. -- Sepia pharaonis Ehrenberg, 1831 [fide Adam and Rees (1966:22)]
TYPE REPOSITORY. -- AMS Holotype C133333 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Howick Island, Queensland, Australia

Sepia incerta Smith, 1916:23, pl 2 fig 6.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:100)]
TYPE REPOSITORY. -- BMNH Syntypes 1890.12.14.43-44 (sepion) [fide Lipinski et al. (2000:103)]
TYPE LOCALITY. -- Port Elizabeth, South Africa [fide Adam and Rees (1966:67)]

Sepia indica D'Orbigny, 1839-1842
In Ferussac and D'Orbigny, 1834-1848:288, Seiche pl 21
[fide Tillier and Boucher-Rodoni (1994:101); taxon dated from plate 25]

CURRENT SYSTEMATIC STATUS. -- Sepia aculeata Van Hasselt, 1835 In Ferussac and D'Orbigny, 1834-1848 [fide Adam and Rees (1966:12)]
TYPE REPOSITORY. -- MNHN Syntypes (3) 1-6-229 and 1-6-230 [fide Lu et al. (1995:318)]
TYPE LOCALITY. -- Bombay, India

Sepia inermis Van Hasselt, 1835
In Ferussac and D'Orbigny, 1834-1848:286, pls 6bis, 20 figs 1-9
[fide Tillier and Boucher-Rodoni (1994:101); taxon dated from plate 6bis]

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepiella inermis [fide Adam and Rees (1966:123)]
TYPE REPOSITORY. -- MNHN Syntypes 1-7-253 to 1-7-259, 1-7-1200 and 1-7-1201 [fide Khromov et al. (1998:134)]
TYPE LOCALITY. -- Bombay, Pondichery, India

Sepia insignis Smith, 1916:25, pl 2 fig 10.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:100)]
TYPE REPOSITORY. -- BMNH Holotype 1920.3.23.5 [fide Lipinski et al. (2000:103)]
TYPE LOCALITY. -- Tongaat Beach, Natal, South Africa

Sepia irvingi Meyer, 1909:333, figs 7-10.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Lu (1998a:169)]
TYPE REPOSITORY. -- Unresolved; Syntypes (4) [fide Meyer (1909:333)]; WAM[?] Type [= Lectotype designation] [fide Adam (1979:139)]; WAM Syntype 4203 [fide Lu (1998a:171); ZMH Syntypes (2) 3842 [fide Guerrero-Kommritz (2001:172)]
TYPE LOCALITY. -- Cockburn Sound, Port Royal and George Sound, Western Australia

Sepia italica Risso, 1854:13, pl 4 figs 1-2.

CURRENT SYSTEMATIC STATUS. -- Sepia elegans Blainville, 1827 [fide Adam and Rees (1966:94)]
TYPE REPOSITORY. -- Unresolved; MHNN?
TYPE LOCALITY. -- Mediterranean

Sepia ivanovi Khromov, 1982:137, figs 1-7.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:101)]
TYPE REPOSITORY. -- ZMMGU Holotype Y-172 [fide Ivanov and Sysoev (2000:108)]
TYPE LOCALITY. -- Mombasa, Zambezi (South-East Africa)

Decorisepia jaenschi Cotton, 1931:41, figs 5-6.

CURRENT SYSTEMATIC STATUS. -- Sepia rex (Iredale, 1926) [fide Lu (1998a:180)]
TYPE REPOSITORY. -- SAMA Holotype D10163 [fide Zeidler and MacPhail (1978:382)]
TYPE LOCALITY. -- Robe, South Australia

Acanthosepion javanicum Rochebrune, 1884b:110.

CURRENT SYSTEMATIC STATUS. -- Sepia aculeata Van Hasselt, 1835 In Ferussac and D'Orbigny, 1834-1848 [fide Adam and Rees (1966:12)]
TYPE REPOSITORY. -- MNHN Holotype 1-6-186 and 1-6-187 (sepion) [fide Lu et al. (1995:318)]
TYPE LOCALITY. -- Java

Sepia (Doratosepion) joubini Massy, 1927:161, pl 18.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:101)]
TYPE REPOSITORY. -- BMNH Syntypes (3) 1926.10.20.38 and 1926.10.20.39-40 [fide Lipinski et al. (2000:103)]
TYPE LOCALITY. -- Tugela River, Cape Natal, South Africa

Sepia jousseaumi Rochebrune, 1884b:117.

CURRENT SYSTEMATIC STATUS. -- Sepia vermiculata Quoy and Gaimard, 1832 [fide Khromov et al. (1998:130)]
TYPE REPOSITORY. -- MNHN Holotype 1-6-221 and 1-6-222 (sepion) [fide Lu et al. (1995:318)]
TYPE LOCALITY. -- Cape of Good Hope

Sepia officinalis jurujubai Oliveira, 1940:29.

CURRENT SYSTEMATIC STATUS. -- Sepioteuthis sepioidea (Blainville, 1823) [fide Adam and Rees (1966:151)]
TYPE REPOSITORY. -- Unresolved
TYPE LOCALITY. -- Jurujuba, Brazil

Sepia kiensis Hoyle, 1885c:194.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Adam and Rees (1966:78)]
TYPE REPOSITORY. -- BMNH Holotype 1889.4.24.80 [fide Lipinski et al. (2000:103)]
TYPE LOCALITY. -- 5∞49'15"S, 132∞14'05"E, off Ki Islands (Araffura Sea)

Sepia kobiensis Hoyle, 1885c:195.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:101)]
TYPE REPOSITORY. -- BMNH Holotype 1889.4.24.81 [fide Lipinski et al. (2000:104)]
TYPE LOCALITY. -- 34∞39'N, 135∞14'E, Bay of Kobe, Japan

Sepia koettlitzi Hoyle and Standen, 1901:1, pl 1.

CURRENT SYSTEMATIC STATUS. -- Sepia pharaonis Ehrenberg, 1831 [fide Adam and Rees (1966:22)]
TYPE REPOSITORY. -- BMNH Holotype 1903.1.2.1 (sepion) [fide Lipinski (2000:104)]
TYPE LOCALITY. -- Zeila, Somaliland

Sepia koilados Reid, 2000:24, figs 7, 14-18.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide; taxon not yet reviewed]
TYPE REPOSITORY. -- NMV Holotype F82459
TYPE LOCALITY. -- NW Shelf, Western Australia

Solitosepia lana Iredale, 1954:66.

CURRENT SYSTEMATIC STATUS. -- Sepia papuensis Hoyle, 1885 [fide Lu (1998a:175)]
TYPE REPOSITORY. -- AMS Holotype C133301 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Low Isles, Queensland, Australia

Sepia latimanus Quoy and Gaimard, 1832:68, pl 2 figs 2-11.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Lu (1998a:171)]
TYPE REPOSITORY. -- MNHN Syntype 1-6-192 [fide Lu et al. (1995:318)]
TYPE LOCALITY. -- Port Dorey, New Guinea

Metasepia pfefferi laxior Iredale, 1926b:240, pl 35 figs 9-10.

CURRENT SYSTEMATIC STATUS. -- Metasepia pfefferi (Hoyle, 1885) [fide Lu (1998a:186)]
TYPE REPOSITORY. -- AMS Holotype C133326 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Masthead Island, Queensland, Australia

Sepia lefebrei D'Orbigny, 1839-1842
In Ferussac and D'Orbigny, 1834-1848:282, Seiche pl 24 figs 1-6
†[fide Tillier and Boucher-Rodoni (1994:101); taxon dated from plate]

CURRENT SYSTEMATIC STATUS. -- Sepia gibba Ehrenberg, 1831 [fide Adam and Rees (1966:100)]
TYPE REPOSITORY. -- MNHN Lectotype 1990 [fide Lu et al. (1995:319)]
TYPE LOCALITY. -- Red Sea

Solitosepia liliana Iredale, 1926a:188, pl 21.

CURRENT SYSTEMATIC STATUS. -- Sepia mestus Gray, 1849 [fide Lu (1998a:171)]
TYPE REPOSITORY. -- AMS Holotype C133300 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Manly Beach, N.S.W., Australia

Arctosepia limata Iredale, 1926a:193, pl 23.

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepia limata [fide Reid (1998:171)]
TYPE REPOSITORY. -- AMS Holotype C133316 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Manly Beach, N.S.W., Australia

Chondrosepia loliginiformis Ruppell and Leuckart, 1828:21, pl 6.

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepioteuthis loliginiformis [fide Ehrenberg (1831:[3])]
TYPE REPOSITORY. -- "Mus. Francof."; SMF Holotype 23805
TYPE LOCALITY. -- Near Mohila, Red Sea

Sepia loligo Linne, 1758:659.

CURRENT SYSTEMATIC STATUS. -- On Official Index of Rejected and Invalid Specific Names in Zoology (ICZN Opinion 1861,† 1997:59)

Sepia loligo Gmelin, 1791:3150.

CURRENT SYSTEMATIC STATUS. -- Todarodes sagittatus (Lamarck, 1798) [fide Adam and Rees (1966:152)]
TYPE REPOSITORY. -- Unresolved
TYPE LOCALITY. -- “Habitat in pelago”

Sepia longipes Sasaki, 1913a:78 pl 3 figs 4-5.

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepia (Doratosepion) longipes [fide Khromov et al. (1998:103)]
TYPE REPOSITORY. -- MSUT[TIU] Syntypes (6) [fide Sasaki (1929:189)]
TYPE LOCALITY. -- Choshi, Kazusa Province, Japan

Sepia (Doratosepion) longipes Sasaki, 1914:619, pl 12 figs 4-5.

CURRENT SYSTEMATIC STATUS. -- Preoccupied by Sepia longipes Sasaki, 1913

Sepia lorigera Wulker, 1910:12, figs 3-4, 11-14.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:104)]
TYPE REPOSITORY. -- ZSM Holotype [specimen not extant?]
TYPE LOCALITY. -- Misaki, Japan

Sepia officinalis lutea Risso, 1854:11.

Nomen nudum [fide Adam and Rees (1966:152)]

Sepia lycidas Gray, 1849:103.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:104)]
TYPE REPOSITORY. -- BMNH Holotype 1963119W [fide Lipinski et al. (2000:104)]
TYPE LOCALITY. -- Canton market, China

Mesembrisepia macandrewi Iredale, 1926a:191, pl 21.

CURRENT SYSTEMATIC STATUS. -- Sepia novaehollandiae Hoyle, 1909 [fide Lu (1998a:174)]
TYPE REPOSITORY. -- AMS Holotype C133328 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Shellharbour, N.S.W., Australia

Glyptosepia macilenta Iredale, 1926a:192, pl 22.

CURRENT SYSTEMATIC STATUS. -- Sepia cultrata Hoyle, 1885 [fide Lu (1998a:166)]
TYPE REPOSITORY. -- AMS Holotype C133305 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Manly Beach, N.S.W., Australia

Sepia madokai Adam, 1939e:77.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:106)]
TYPE REPOSITORY. -- MSUT Holotype [fide Khromov et al. (1998:106)]
TYPE LOCALITY. -- Tokyo Bay, Japan

Sepiadarium malayense Robson, 1932b:28, figs 3-4.

CURRENT SYSTEMATIC STATUS. -- Sepiadarium kochi Steenstrup, 1881 [fide Voss (1963b:38)]
TYPE REPOSITORY. -- BMNH Holotype 1938.5.9.42 [fide Lipinski et al. (2000:104)]
TYPE LOCALITY. -- Singapore fish market

Sepia mammalita Leach, 1835
In Ferussac and D'Orbigny, 1834-1848:277, Seiche pl 4bis
[fide Tillier and Boucher-Rodoni (1994:101); taxon dated from plate]

CURRENT SYSTEMATIC STATUS. -- ?Sepia tuberculata Lamarck, 1798 [fide Adam and Rees (1966:106)]
TYPE REPOSITORY. -- MNHN Type; specimen not extant [fide Lu et al. (1995:320)]
TYPE LOCALITY. -- Not designated

Sepiella mangkangunga Reid and Lu, 1998:74, figs 1-11.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide; taxon not yet reviewed]
TYPE REPOSITORY. -- NTM Holotype P10606
TYPE LOCALITY. -- Northern Territory, Australia

Sepia marmorata Ruppell In Wulker, 1920:54, 55.

Nomen nudum.

Sepia (Doratosepion) mascarensis Filippova and Khromov, 1991:63, pl 1.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:106)]
TYPE REPOSITORY. -- ZMMGU Holotype Y-349 [fide Ivanov and Sysoev (2000:109)]
TYPE LOCALITY. -- Saya-de-Malha Bank (Indian Ocean)

Sepia media Barbut, 1783:75, pl 8 fig 3.

CURRENT SYSTEMATIC STATUS. -- Todarodes sagittatus (Lamarck, 1798) [fide Adam and Rees (1966:152)]
TYPE REPOSITORY. -- Unresolved
TYPE LOCALITY. -- “It inhabits the seas”

Sepia media Linne, 1758:659.

CURRENT SYSTEMATIC STATUS. -- Valid species; Loligo (Alloteuthis) media [fide Vecchione et al. (1998:220)]
TYPE REPOSITORY. -- Unresolved; LS?
TYPE LOCALITY. -- “Pelago”

Sepia officinalis mediterranea Ninni, 1884:169, pl 2 fig 2.

CURRENT SYSTEMATIC STATUS. -- Sepia officinalis Linne, 1758 [fide Khromov et al. (1998:81)]
TYPE REPOSITORY. -- Unresolved
TYPE LOCALITY. -- Adriatic

Sepia (Acanthosepia) mercatoris Adam, 1937:56, figs 6-10.

CURRENT SYSTEMATIC STATUS. -- Sepia bertheloti D'Orbigny, 1835 [fide Adam and Rees (1966:57)]
TYPE REPOSITORY. -- Unresolved; IRSNB?
TYPE LOCALITY. -- Rufisque, Senegal

Sepia mestus Gray, 1849:108.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Lu (1998a:171)]
TYPE REPOSITORY. -- BMNH Syntypes 1842.11.9.84 (sepion) and 1842.11.11.13 (sepion) [fide Lipinski et al. (2000:104)]
TYPE LOCALITY. -- Australia

Sepia microcheirus Gray, 1849:107.

CURRENT SYSTEMATIC STATUS. -- Sepiella inermis (Van Hasselt, 1835 In Ferussac and D'Orbigny, 1834-1848) [fide Adam and Rees (1966:124)]
TYPE REPOSITORY. -- Unresolved; BMNH Syntypes [?= 1963121, 1963123, 1963124], not extant [fide Lipinski et al. (2000:114)]
TYPE LOCALITY. -- India

Sepia micocrosmus Oken, 1815:344.

CURRENT SYSTEMATIC STATUS. -- Undetermined
TYPE REPOSITORY. -- Unresolved
TYPE LOCALITY. -- Unresolved

Sepia microcotyledon Ortmann, 1891:673, pl 46 fig 1a-c.

CURRENT SYSTEMATIC STATUS. -- Sepia aculeata Van Hasselt, 1835 In Ferussac and D'Orbigny, 1834-1848 [fide Adam and Rees (1966:12)]
TYPE REPOSITORY. -- MZUS Holotype [fide Johnson (1977:238)]
TYPE LOCALITY. -- Ceylon [Sri Lanka]

Sepia minima Lesueur, 1821:100.

Nomen nudum [fide Adam and Rees (1966:152)]; New name for "Sepia sepiola. Peron"

Sepia minor Seba, 1759 In 1734-1765:pl 3.
[fide Adam and Rees (1966:152)]

CURRENT SYSTEMATIC STATUS. -- Invalid; non-binomial names [fide Sherborn (1902:xlix)]

Tenuisepia mira Cotton, 1932:546, figs 7-9.

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepia mira [fide Reid (1998:160)]
TYPE REPOSITORY. -- SAMA Holotype D10507 (sepion) [fide Zeidler and MacPhail (1978:382)]
TYPE LOCALITY. -- North-West Islet, Queensland, Australia [fide Adam and Rees (1966:87)]

Sepia (Doratosepion) mirabilis Khromov, 1988:785.

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepia mirabilis [fide Khromov et al. (1998:109)]
TYPE REPOSITORY. -- ZMMGU Holotype Y-300 [fide Ivanov and Sysoev (2000:108)]
TYPE LOCALITY. -- Sokotra Island (West Arabian Sea)

Sepia misakiensis Wulker, 1910:15, figs 5-6, 19-22.

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepia (Doratosepion) misakiensis [fide Kubodera (2000a:1053)]
TYPE REPOSITORY. -- Unresolved; ZSM?
TYPE LOCALITY. -- Misaki, Japan

Sepia mozambica Rochebrune, 1884b:118.

CURRENT SYSTEMATIC STATUS. -- Sepia latimanus Quoy and Gaimard, 1832 [fide Adam and Rees (1966:33)]
TYPE REPOSITORY. -- MNHN Holotype 1-5-125 (sepion) [fide Lu et al. (1995:321)]
TYPE LOCALITY. -- Detroit of Mozambique [fide Lu et al. (1995:321)]

Sepia mucronata Rafinesque, 1814:29.

Nomen nudum [fide Adam and Rees (1966:152)]

Sepia murrayi Adam and Rees, 1966:63, text-fig 260, pls 18, 43.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:109)]
TYPE REPOSITORY. -- BMNH Holotype 196381W [fide Lipinski et al. (2000:104)]
TYPE LOCALITY. -- 25∞35'-25∞43'N, 56∞42'-56∞39'E, Gulf of Oman

Sepia myrsus Gray, 1849:108.

CURRENT SYSTEMATIC STATUS. --?Sepia simoniana Thiele, 1920 [fide Adam and Rees (1966:112)]
TYPE REPOSITORY. -- BMNH Holotype 1841.4.6.89 (sepion) [fide Lipinski et al. (2000:104)]
China?

Sepia nanshiensis Li and Chen, 1989:6-12, figs 1-4.

CURRENT SYSTEMATIC STATUS. -- Sepia vietnamica Khromov, 1987 [fide Khromov et al. (1998:130)]
TYPE REPOSITORY. -- SCSIO Holotype[?] AS-M20
TYPE LOCALITY. -- South China Sea

Sepia natalensis Massy, 1925:212, pls 11, 14.

CURRENT SYSTEMATIC STATUS. -- Sepia simoniana Thiele, 1920 [fide Voss (1962d:247)]
TYPE REPOSITORY. -- Unresolved; NMP?
TYPE LOCALITY. -- NE of Bird Island

Sepia nigra Bosc, 1802:47.

CURRENT SYSTEMATIC STATUS. --?Dosidicus gigas (D'Orbigny, 1835 In 1834-1847) [fide Adam and Rees (1966:152)]
TYPE REPOSITORY. -- Unresolved; “Sepia tunicata Gmel. Syst. Nat.”
TYPE LOCALITY. -- “Se trouve dans les mers da Chile”

Sepia novaehollandiae Hoyle, 1909:266.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Lu (1998a:174)]
TYPE REPOSITORY. -- MNHN Syntype 1-6-194 (sepion) [fide Lu et al. (1995:321)]
TYPE LOCALITY. -- Kangaroo Island, S. Australia

Solitosepia occidua Cotton, 1929:88, pl 14.

CURRENT SYSTEMATIC STATUS. -- Sepia papuensis Hoyle, 1885 [fide Lu (1998a:175)]
TYPE REPOSITORY. -- SAMA Holotype D13627 [fide Zeidler and MacPhail (1978:382)]
TYPE LOCALITY. -- Rottnest Island, Western Australia

Sepia octopoda Peron In Lesueur, 1821:101.

Nomen nudum [fide Lu et al. (1995:321)]

Sepia octopodia Linne, 1758:658.

CURRENT SYSTEMATIC STATUS. -- On Official Index of Rejected and Invalid Specific Names in Zoology (ICZN Opinion 1861, 1997:59)

Sepia octopodia Molina, 1789:173.

CURRENT SYSTEMATIC STATUS. -- Preoccupied by Sepia octopodia Linne, 1758.

Sepia octopus Gmelin, 1791:3149.

CURRENT SYSTEMATIC STATUS. -- On Official Index of Rejected and Invalid Specific Names in Zoology (ICZN Opinion 1861, 1997:59)

Acanthosepion oculiferum Rochebrune, 1884b:107.

CURRENT SYSTEMATIC STATUS. -- Sepia hierredda Rang, 1835 In Ferussac and D'Orbigny, 1834-1848 [fide Khromov et al. (1998:99)]
TYPE REPOSITORY. -- MNHN Holotype 1-6-195 [fide Lu et al. (1995:321)]
TYPE LOCALITY. -- Goree, Senegal [Gulf of Guinea]

Sepia officinalis Linne, 1758:658.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:81)]
TYPE REPOSITORY. -- Unresolved; LS?
TYPE LOCALITY. -- “Oceano”

Sepia omani Adam and Rees, 1966:92, pls 22, 41.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:110)]
TYPE REPOSITORY. -- BMNH Holotype 1963107W [fide Lipinski et al. (2000:104)]
TYPE LOCALITY. -- Gulf of Oman, 25∞10'48"-25∞09'48"N, 56∞47'30"E

Glyptosepia opipara Iredale, 1926a:191, pl 22 figs 7-8.

CURRENT SYSTEMATIC STATUS. -- Valid species Sepia opipara [fide Lu (1998a:175)]
TYPE REPOSITORY. -- AMS Holotype C133330 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Masthead Island, Capricorn Group, Queensland, Australia

Sepia orbignyana Ferussac In D'Orbigny, 1826:156.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:110)]
TYPE REPOSITORY. -- MNHN Holotype 1-4-88 (sepion) [fide Lu et al. (1995:322)]
TYPE LOCALITY. -- La Rochelle, France

Sepia ornata Rang, 1837:76, pl 101.

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepiella ornata [fide Adam and Rees (1966:127)]
TYPE REPOSITORY. -- MNHN Syntypes 1-7-231 (sepion), 1-7-232, 1-7-233 (sepion), 1-7-234 [fide Lu et al. (1995:322)]
TYPE LOCALITY. -- Coast of Goree, Senegal [Gulf of Guinea]

Mesembrisepia ostanes Iredale, 1954:69, pl 4 figs 5-6.

CURRENT SYSTEMATIC STATUS. -- Sepia novaehollandiae Hoyle, 1909 [fide Lu (1998a:174)]
TYPE REPOSITORY. -- AMS Holotype C133311 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Stanley, Tasmania

Sepiella ovata Pfeffer, 1884:14, figs 18-18a.

CURRENT SYSTEMATIC STATUS. -- ?Sepiella japonica Sasaki, 1929 [fide Adam and Rees (1966:128)]
TYPE REPOSITORY. -- Unresolved; ZMH Holotype [species not traced in Guerrero-Kommritz (2001)]; IRSNB Type [fide F.G. Hochberg (pers. comm.)]
TYPE LOCALITY. -- Not designated

Sepia pagenstecheri Pfeffer, 1884:9, figs 12-12a.

CURRENT SYSTEMATIC STATUS. -- ?Sepia recurvirostra Steenstrup, 1875 [fide Adam and Rees (1966:28)]
TYPE REPOSITORY. -- Unresolved; ZMH Holotype [species not traced in Guerrero-Kommritz (2001)]; IRSNB Type [fide F.G. Hochberg (pers. comm.)]
TYPE LOCALITY. -- Hong Kong

Acanthosepion pageorum Iredale, 1954:76, pl 4 figs 7-9.

CURRENT SYSTEMATIC STATUS. -- Sepia smithi Hoyle, 1885 [fide Lu (1998a:184)]
TYPE REPOSITORY. -- AMS Holotype C133315 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Keppel Bay, Queensland, Australia

Sepia palmata Owen, 1881:134, pl 24-25.

CURRENT SYSTEMATIC STATUS. -- Sepia apama Gray, 1849 [fide Lu (1998a:160)]
TYPE REPOSITORY. -- "Type lost" [fide Lu (1998a)]
TYPE LOCALITY. -- Off the shore of Norfolk Island, Australia

Sepia papillata Quoy and Gaimard, 1832:61, pl 1 figs 6-14.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:111)]
TYPE REPOSITORY. -- MNHN Syntype 1-4-78 [fide Lu et al. (1995:322)]
TYPE LOCALITY. -- Cape of Good Hope [fide Adam and Rees (1966:109)]

Sepia papuensis Hoyle, 1885c:197.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Lu (1998a:175)]
TYPE REPOSITORY. -- BMNH Syntype 1889.4.24.68 [fide Lipinski et al. (2000:104)]; NMGW Syntype
TYPE LOCALITY. -- 9∞59'S, 139∞42'E (Arafura Sea)

Sepia (Doratosepion) pardalis Sasaki, 1914:614, pl 12 figs 1-3.

CURRENT SYSTEMATIC STATUS. -- Sepia pardex Sasaki, 1913 [fide Kubodera (2000a:1051)] [change in species name in English translation of Sepia pardex Sasaki, 1913, published in Japanese]
TYPE REPOSITORY. -- MSUT Holotype [fide Khromov et al. (1998:111)]
TYPE LOCALITY. -- Kajiyama, Awa Province, Japan

Sepia pardex Sasaki, 1913a:74.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Kubodera (2000a:1051)]
TYPE REPOSITORY. -- MSUT Holotype [fide Khromov et al. (1998:111)]
TYPE LOCALITY. -- Kajiyama, Awa Province, Japan

Amplisepia parysatis Iredale, 1954:71, pl 4 figs 1-2.

CURRENT SYSTEMATIC STATUS. -- Sepia apama Gray, 1849 [fide Lu (1998a:160)]
TYPE REPOSITORY. -- AMS Holotype C133307 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Timor Sea

Sepia pelagica Bosc, 1802:46 (vol 1), pl 1 figs 1-2.

CURRENT SYSTEMATIC STATUS. -- Valid species; Hyaloteuthis pelagica [fide Gray (1849:63)]
TYPE REPOSITORY. -- Unresolved [fide Wormuth (1998:379)]
TYPE LOCALITY. -- “Trouvee en pleine mer, par Bosc dans l’estomac d’une dorade”

Solitosepia rozella peregrina Iredale, 1926b:238.

CURRENT SYSTEMATIC STATUS. -- Sepia rozella Iredale, 1926 [fide Lu (1998a:181)]
TYPE REPOSITORY. -- AMS Holotype C133322 [fide Rudman (1983:68)]
TYPE LOCALITY. -- North-west Island, Queensland, Australia

Sepia peronii Lesueur, 1821:101.

Nomen nudum; Octopod [fide Robson (1929d:216)]; New name for “Sepia octopa. Peron”

Sepia peterseni Appellof, 1886:23, pl 2 figs 1-6, pl 3 fig 21.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:112)]
TYPE REPOSITORY. -- ZMUU Syntypes 148a-b [fide Wallin (1991:66)]
TYPE LOCALITY. -- Japan

Sepia (Metasepia) pfefferi Hoyle, 1885c:199.

CURRENT SYSTEMATIC STATUS. -- Valid species; Metasepia pfefferi [fide Lu (1998a:186)]
TYPE REPOSITORY. -- BMNH Holotype 1890.1.24.11 [fide Lipinski et al. (2000:104)]
TYPE LOCALITY. -- 9∞59'S, 139∞42'E (Arafura Sea)

Sepia pharaonis Ehrenberg, 1831:[4].

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:112)]
TYPE REPOSITORY. -- ZMB Lectotype Moll.-000083 [fide Glaubrecht and Salcedo-Vargas (2000:277)]
TYPE LOCALITY. -- Gulf of Suez, Red Sea [fide Glaubrecht and Salcedo-Vargas (2000:277)]

Sepia plana Lu and Reid, 1997:279, figs 2-9.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide; taxon not yet reviewed]
TYPE REPOSITORY. -- NMV Holotype F77206
TYPE LOCALITY. -- 14∞S, 121∞E, Western Australia

Sepia plangon Gray, 1849:104.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Lu (1998a:178)]
TYPE REPOSITORY. -- BMNH Syntypes 1963126W and 1842.11.9.85 (sepion) [fide Lipinski et al. (2000:104)]
TYPE LOCALITY. -- Port Jackson; Australia

Sepia (Sepia) plathyconchalis Filippova and Khromov, 1991:66, pl 2.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:112)]
TYPE REPOSITORY. -- ZMMGU Holotype Y-351 [fide Ivanov and Sysoev (2000:109)]
TYPE LOCALITY. -- Saya-de-Malha Bank (Indian Ocean)

Sepia polynesica Pfeffer, 1884:11, figs 14-14a.

CURRENT SYSTEMATIC STATUS. --?Sepia plangon Gray, 1849 [fide Adam and Rees (1966:44)]
TYPE REPOSITORY. -- ZMH Holotype [species not traced in Guerrero-Kommritz (2001)]
TYPE LOCALITY. – “Sudsee”

Sepia prashadi Winckworth, 1936:17, pl 4.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:117)]
TYPE REPOSITORY. -- IMC Holotype [fide Khromov et al. (1998:117)]
TYPE LOCALITY. -- Beach, Madras, India

Sepia prionota Voss, 1962a:169.

CURRENT SYSTEMATIC STATUS. -- Sepia papuensis Hoyle, 1885 [fide Adam and Rees (1966:41)]
TYPE REPOSITORY. -- NMNH Holotype 575326 [fide Roper and Sweeney (1978:11)]
TYPE LOCALITY. -- Off Sirun Island, Sulu Archipelago, Tawi Tawi group, Philippine Islands

Sepia pulchra Roeleveld and Liltved, 1985:2, figs 1-14.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:117)]
TYPE REPOSITORY. -- SAM Holotype S-1036 [fide Khromov et al. (1998:117)]
TYPE LOCALITY. -- Cape Peninsula, Llandudno, South Africa

Sepia ramani Neethiselvan, 2001:82, figs 1, 3-4, 6 [sic].

CURRENT SYSTEMATIC STATUS. -- Valid species [fide; taxon not yet reviewed]
TYPE REPOSITORY. -- FCRIRM Holotype LA3
TYPE LOCALITY. -- Tuticorin, SE coast of India

Sepia rappiana Ferussac, 1835
In Ferussac and D'Orbigny, 1834-1848:283, Seiche pl 10
[fide Tillier and Boucher-Rodoni (1994:101); authorship and taxon dated from plate]

CURRENT SYSTEMATIC STATUS. -- Sepia latimanus Quoy and Gaimard, 1832 [fide Adam and Rees (1966:33)]
TYPE REPOSITORY. -- MNHN Type; specimen not extant [fide Lu et al. (1995:323)]
TYPE LOCALITY. -- Not designated

Sepia recurvirostra Steenstrup, 1875:475.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:118)]
TYPE REPOSITORY. -- ZMUC Lectotype [fide Adam (1965)]
TYPE LOCALITY. -- South China Sea [fide Kristensen and Knudsen (1983:223)]

Sepia reesi Adam, 1979:200, pl 4 fig 3.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Lu (1998a:180)]
TYPE REPOSITORY. -- WAM Holotype 497.76 (sepion) [fide Slack-Smith (1983:75)]
TYPE LOCALITY. -- Rottnest Island, Western Australia

Decorisepia rex Iredale, 1926a:193, pl 22 figs 9-10.

CURRENT SYSTEMATIC STATUS. -- Sepia hedleyi Berry, 1918 [fide Reid (2001b:83)]
TYPE REPOSITORY. -- AMS Holotype C127593 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Manly Beach, N.S.W., Australia [fide Lu (1998a:180)]

Arctosepia rhoda Iredale, 1954:75, pl 4, figs 10-12.

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepia rhoda [fide Reid (2000:39)]
TYPE REPOSITORY. -- AMS Holotype C133319 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Point Cloates, Western Australia (22o43'S, 113o40'E) [fide Reid (2000)]

Rhombosepion robsoni Massy, 1927:159, pl 17.

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepia robsoni [fide Voss (1962d:248)]
TYPE REPOSITORY. -- BMNH Holotype 1926.10.20.8 [fide Lipinski et al. (2000:104)]
TYPE LOCALITY. -- Hout Bay, South Africa [fide Khromov et al. (1998:118)]

Sepia robsoni Sasaki, 1929:179, pl 17 figs 2-3.

CURRENT SYSTEMATIC STATUS. -- Junior homonym of Sepia robsoni (Massy, 1927)
TYPE REPOSITORY. -- MSUT[TIU] Syntype(s)
TYPE LOCALITY. -- Tokyo Bay, Japan

Sepia rostrata D'Orbigny, 1835
In Ferussac and D'Orbigny, 1834-1848:284, Seiche pl 8 fig 6 and pl 24
[fide Tillier and Boucher-Rodoni (1994:101); taxon dated from plate 8]

CURRENT SYSTEMATIC STATUS. -- Undetermined; [see Adam and Rees (1966:154)]
TYPE REPOSITORY. -- MNHN Syntypes 1-4-114, 1-4-115 (sepion), 1-4-1924, 1-4-119 [fide Lu et al. (1995:324)]
TYPE LOCALITY. -- Bombay, India

Sepia rouxii D'Orbigny, 1839-1842
In Ferussac and D'Orbigny, 1834-1848:271, Seiche pl 19
[fide Tillier and Boucher-Rodoni (1994:101); taxon dated from plate]

CURRENT SYSTEMATIC STATUS. -- Sepia pharaonis Ehrenberg, 1831 [fide Adam and Rees (1966:22)]
TYPE REPOSITORY. -- MNHN Syntypes 1-4-95 + 1-4-96 (sepion), 1-4-105, 1-4-94, 1-5-150, 1-5-151, 1-5-151bis [fide Lu et al. (1995:324)]
TYPE LOCALITY. -- Bombay, India and Red Sea

Solitosepia rozella Iredale, 1926a:190, pl 21 figs 6-7.

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepia rozella [fide Lu (1998a:181)]
TYPE REPOSITORY. -- AMS Holotype C133336 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Manly Beach, N.S.W., Australia

Sepia rubens Philippi, 1844:203.

CURRENT SYSTEMATIC STATUS. -- Sepia orbignyana Ferussac In D'Orbigny, 1826 [fide Adam and Rees (1966:87)]
TYPE REPOSITORY. -- Unresolved
TYPE LOCALITY. -- Naples

Sepia rugosa Bosc, 1792:24, pl 5 figs 1-2.

CURRENT SYSTEMATIC STATUS. -- Octopus vulgaris Cuvier, 1797 [fide Mangold (1998:525)]
TYPE REPOSITORY. -- Unresolved
TYPE LOCALITY. -- Senegal [?]

Sepia rugosa Bowdich, 1822:pl 1 fig 1.

Nomen nudum; Sepia officinalis Linne, 1758 [fide Adam and Reese (1966:154)]

Sepia rupellaria D'Orbigny, 1834
In Ferussac and D'Orbigny, 1834-1848]:274, Seiche pl 3 figs 10-13
[fide Tillier and Boucher-Rodoni (1994:101); taxon dated from plate]

CURRENT SYSTEMATIC STATUS. -- Sepia elegans Blainville, 1827 [fide Adam and Rees (1966:94)]
TYPE REPOSITORY. -- MNHN Syntypes (2) 1-6-200 (sepions) [fide Lu et al. (1995:324)]
TYPE LOCALITY. -- La Rochelle, France

Sepia kobiensis sagamiana Sasaki, 1929:207, text-figs 112-114.

Nomen nudum; illustrations for Sepia kobiensis andreanoides Hoyle, 1885

Sepia (Doratosepion) sasakii Wakiya and Ishikawa, 1921:290, figs 11a-c.

CURRENT SYSTEMATIC STATUS. --?Sepia andreana Steenstrup, 1875 [fide Nesis (1987a:114)]
TYPE REPOSITORY. -- Unresolved; Syntypes (19)
TYPE LOCALITY. -- Tokyo market (2); Miyagi Prefecture (16); Aomori Prefecture (1)

Sepia savignyi Blainville, 1827:285.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:119)]
TYPE REPOSITORY. -- MNHN Neotype 1-6-188bis and 1-6-188 (sepion) [fide Rochebrune (1884b)]
TYPE LOCALITY. -- Tor, Red Sea [fide Lu et al. (1995:324)]

Sepia (Doratosepion) saya Khromov, Nikitina and Nesis, 1991:13, pl 1.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:119)]
TYPE REPOSITORY. -- ZMMGU Holotye Y-357 [fide Ivanov and Sysoev (2000:108)]
TYPE LOCALITY. -- Saya-de-Malha Bank (Indian Ocean)

Sepia senta Lu and Reid, 1997:286, figs 9-16.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide; taxon not yet reviewed]
TYPE REPOSITORY. -- NMV Holotype F77200
TYPE LOCALITY. -- 14∞S, 122∞E, Western Australia

Octopodia sepia Schneider, 1784:109.

CURRENT SYSTEMATIC STATUS. -- Rejected specific name [fide Opinion 233, ICZN (1954:278)]

Sepia sepiola Linne, 1758:659.

CURRENT SYSTEMATIC STATUS. -- Sepiola rondeletii Leach, 1834 In Ferussac and D'Orbigny, 1834-1848 [see Hoyle (1910b:412)]
TYPE REPOSITORY. -- Unresolved; LS?
TYPE LOCALITY. -- Mediterranean

Sepia sewelli Adam and Rees, 1966:61, text-fig 273, pls 17, 46.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:119)]
TYPE REPOSITORY. -- BMNH Holotype 196378W [fide Lipinski et al. (2000:105)]
TYPE LOCALITY. -- 11∞57'12"-11∞56'42"N, 50∞35'00"-50∞39'02"E, near Cape Guardafui, Zanzibar [Somalia]

Sepia simoniana Thiele, 1920:436, pl 52 figs 5-13.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:121)]
TYPE REPOSITORY. -- ZMB Lectotype Moll.-110030 [fide Glaubrecht and Salcedo-Vargas (2000:278)]
TYPE LOCALITY. -- Simio's Bay (South Africa)

Sepia sinensis D'Orbigny, 1835
In Ferussac and D'Orbigny, 1834-1848:289, Seiche pl 9 figs 1-2
[fide Tillier and Boucher-Rodoni (1994:101); taxon dated from plate]

CURRENT SYSTEMATIC STATUS. -- ?Sepiella japonica Sasaki, 1929 [fide Adam and Rees (1966:128)]
TYPE REPOSITORY. -- Type specimen not designated [see also Lu et al. (1995:325)]
TYPE LOCALITY. -- Not designated

Sepia singalensis Goodrich, 1896:3, pl 1 figs 4-8.

CURRENT SYSTEMATIC STATUS. -- Sepia pharaonis Ehrenberg, 1831 [fide Adam and Rees (1966:22)]
TYPE REPOSITORY. -- Unresolved; ZSI? Syntypes (2)
TYPE LOCALITY. -- Columbo and Point Galle(?)

Sepia singaporensis Pfeffer, 1884:10, figs 13-13a.

CURRENT SYSTEMATIC STATUS. -- Sepia recurvirostra Steenstrup, 1875 [fide Adam and Rees (1966:28)]
TYPE REPOSITORY. -- Museum Godeffroy [fide Pfeffer (1884:11)]; IRSNB Type [fide F.G. Hochberg (pers. comm.)]
TYPE LOCALITY. -- Singapore

Sepia sinope Gray, 1849:106.

CURRENT SYSTEMATIC STATUS. -- Sepia australis Quoy and Gaimard, 1832 [fide Khromov et al. (1998:86)]
TYPE REPOSITORY. -- BMNH Holotype 196385W [fide Lipinski et al. (2000:105)]
TYPE LOCALITY. -- China?

Sepia smithi Hoyle, 1885c:190.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Lu (1998a:184)]
TYPE REPOSITORY. -- Unresolved; BMNH Syntypes (2) 1889.4.24.66-67 [fide Lipinski et al. (2000:105)]; NMGW? Syntyypes (2?)
TYPE LOCALITY. -- 9∞59'S, 139∞42'E (Arafura Sea)

Sepia (Doratosepion) sokotriensis Khromov, 1988:788.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:122)]
TYPE REPOSITORY. -- ZMMGU Holotype Y-302 [fide Ivanov and Sysoev (2000:108)]
TYPE LOCALITY. -- Sokotra Island (West Arabian Sea)

Acanthosepion spinigerum Rochebrune, 1884b:103.

CURRENT SYSTEMATIC STATUS. -- Sepia rostrata D'Orbigny, 1835 In Ferussac and D'Orbigny, 1834-1848 and Sepia brevimana Steenstrup, 1875 [fide Adam and Rees (1966:5)]
TYPE REPOSITORY. -- MNHN Holotype 1-4-118 plus 1-4-116 (sepion) [fide Lu et al. (1995:325)]
TYPE LOCALITY. -- Trinquemale, Indian Sea

Sepia stellifera Homenko and Khromov, 1984:1150, figs 1-9.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:122)]
TYPE REPOSITORY. -- ZMMGU Holotype Y-260 [fide Ivanov and Sysoev (2000:109)]
TYPE LOCALITY. -- East Arabian Sea

Sepia subaculeata Sasaki, 1913a:73, pl 3 fig 6a,b.

CURRENT SYSTEMATIC STATUS. -- Sepia lycidas Gray, 1849 [fide Khromov et al. (1998:104)]
TYPE REPOSITORY. -- MSUT[TIU] Syntypes [fide Sasaki (1929:175)]
TYPE LOCALITY. -- Tokyo fish market

Solitosepia submestus Iredale, 1926b:238, pl 35 figs 3-4.

CURRENT SYSTEMATIC STATUS. -- Sepia papuensis Hoyle, 1885 [fide Lu (1998a:175)]
TYPE REPOSITORY. -- AMS Holotype C133325 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Masthead Island, Queensland, Australia

Sepia subplana Lu and Boucher-Rodoni, 2001:370, figs 1-13.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide; taxon not yet reviewed]
TYPE REPOSITORY. -- MNHN Holotype 2213
TYPE LOCALITY. -- Wallis and Futuna Islands (11∞54’S, 179∞32’W)

Sepia (Doratosepion) subtenuipes Okutani and Horikawa, 1987:57, 2 figs.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:122)]
TYPE REPOSITORY. -- NSMT Holotype Mo-64227 [fide Kubodera and Tsuchiya (1993:unpaginated)]
TYPE LOCALITY. -- Tosa Bay, Japan

Sepia subulata Bosc, 1802:46.

CURRENT SYSTEMATIC STATUS. --?Alloteuthis subulata (Lamarck, 1798) [fide Adam and Rees (1966:154)]
TYPE REPOSITORY. -- Unresolved
TYPE LOCALITY. -- Not designated

Sepia sulcata Hoyle, 1885c:192.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:122)]
TYPE REPOSITORY. -- BMNH Holotype 1889.4.24.77 [fide Lipinski et al. (2000:105)]
TYPE LOCALITY. -- 5∞49'15"S, 132∞14'15"E, off Kii Islands, Arafura Sea

Sepia (Doratopsepion) tala Khromov, Nikitina and Nesis, 1991:17, pl 2.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:125)]
TYPE REPOSITORY. -- ZMMGU Holotype Y-359 [fide Ivanov and Sysoev (2000:108)]
TYPE LOCALITY. -- Off Cape Tala, Southwest Madagascar

Sepia tanybracheia Reid, 2000:47, figs 7, 31-36.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide; taxon not yet reviewed]
TYPE REPOSITORY. -- NMV Holotype F82479
TYPE LOCALITY. -- Ashmore Islands, Western Australia

Sepia tenuipes Sasaki, 1929:193, pl 18 figs 1-9.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:125)]
TYPE REPOSITORY. -- MSUT[TIU] Holotype [fide Khromov et al. (1998:125)]
TYPE LOCALITY. -- Ibaraki Prefecture, Japan

Sepia thurstoni Adam and Rees, 1966:2, pls 1 figs 1-4, 41 fig 235.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:126)]
TYPE REPOSITORY. -- BMNH Holotype 1894.9.6.2-4 [fide Lipinski et al. (2000:105)]
TYPE LOCALITY. -- Rameswaram Island, Madras, India

Sepia tigris Sasaki, 1929:168, text-fig 167, pl 28 figs 13-16.

CURRENT SYSTEMATIC STATUS. -- Sepia pharaonis Ehrenberg, 1831 [fide Adam and Rees (1966:22)]
TYPE REPOSITORY. -- Unresolved [fide Taki and Igarashi (1967:1)]; FMT?
TYPE LOCALITY. -- Taihoku fish market, Formosa [Taiwan]

Sepia tokioensis Ortmann, 1888:653, pl 23 fig 3.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:127)]
TYPE REPOSITORY. -- MZUS Syntypes [fide Johnson (1977:239)]
TYPE LOCALITY. -- Tokyo Bay, Japan

Sepia torosa Ortmann, 1888:652, pl 23 fig 2.

CURRENT SYSTEMATIC STATUS. -- Sepia pharaonis Ehrenberg, 1831 [fide Adam and Rees (1966:22)]
TYPE REPOSITORY. -- MZUS Holotype [fide Johnson (1977:239)]
TYPE LOCALITY. -- Tokyo Bay, Japan

Sepia tourannensis Souleyet, 1852:33, pl 3 figs 6-12.

CURRENT SYSTEMATIC STATUS. -- Sepiella inermis (Van Hasselt, 1835 In Ferussac and D'Orbigny, 1834-1848) [fide Adam and Rees (1966:124)]
TYPE REPOSITORY. -- MNHN Syntype 1-7-248 and 1-7-249 [fide Lu et al. (1995:326)]
TYPE LOCALITY. -- Bay of Touranne, Cochin China [Vietnam]

Sepia kobiensis toyamensis Sasaki, 1929:209, text-fig 115, pl 19 figs 8-12.

CURRENT SYSTEMATIC STATUS. -- Sepia kobiensis Hoyle, 1885 [fide Khromov et al. (1998:101)]
TYPE REPOSITORY. -- FMHU Syntypes (5)
TYPE LOCALITY. -- Toyama Bay, Japan

Arctosepia treba Iredale, 1954:75.

CURRENT SYSTEMATIC STATUS. -- Sepia braggi Verco, 1907 [fide Lu (1998a:161)]
TYPE REPOSITORY. -- AMS Syntypes (2) C133324 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Stanley, Tasmania

Doratosepion trygoninum Rochebrune, 1884b:97, pl 5 fig 1.

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepia trygonina [fide Adam and Rees (1966:84)]
TYPE REPOSITORY. -- MNHN Type; specimen not extant [fide Lu et al. (1995:326)]
TYPE LOCALITY. -- Red Sea

Sepia tuberculata Lamarck, 1798:130.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:128)]
TYPE REPOSITORY. -- MNHN Syntype 1-4-83 (sepion) [fide Lu et al. (1995:326)]
TYPE LOCALITY. -- "Inhabits the Indian sea"

Sepia tullbergi Appellof, 1886:26, pl 2 figs 7-14.

CURRENT SYSTEMATIC STATUS. -- Valid species; Metasepia tullbergi [fide† Nateewathana (1998:323)]
TYPE REPOSITORY. -- ZMUU Holotype 295 [fide Wallin (1991:66)]
TYPE LOCALITY. -- Nagasaki, Japan

Sepia tunicata Molina, 1782:199.

CURRENT SYSTEMATIC STATUS. -- Dosidicus gigas (D'Orbigny, 1835 In 1834-1847) [fide Adam and Rees (1966:154)]
TYPE REPOSITORY. -- Unresolved
TYPE LOCALITY. -- Not designated [Chile]

Hemisepius typicus Steenstrup, 1875:472, pls 1-2.

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepia typica [fide Khromov et al. (1998:129)]
TYPE REPOSITORY. -- ZMUC Holotype [fide Kristensen and Knudsen (1983:224)]
TYPE LOCALITY. -- Off Table Bay, South Africa

Sepia unguiculata Molina, 1782:199.

CURRENT SYSTEMATIC STATUS. -- Taningia danae Joubin, 1931 [fide Clarke (1966:190)]
TYPE REPOSITORY. -- Unresolved
TYPE LOCALITY. -- Not designated [Chile]

Crumenasepia ursulae Cotton, 1929:90, pl 15 fig 3-4.

CURRENT SYSTEMATIC STATUS. -- Sepia pharaonis Ehrenberg, 1831 [fide Adam and Rees (1966:22)]
TYPE REPOSITORY. -- SAMA Holotype D10013 (sepion) [fide Zeidler and MacPhail (1978:381)]
TYPE LOCALITY. -- Cottesloe, Western Australia

Sepia varietas Peron In Lesueur, 1821:101.

Nomen nudum [fide Adam and Rees (1966:155)]

Sepia venusta Pfeffer, 1884:12, figs 15-15a.

CURRENT SYSTEMATIC STATUS. --?Sepia pharaonis Ehrenberg, 1831 [fide Adam and Rees (1966:22)]
TYPE REPOSITORY. -- ZMH Holotype (sepion missing) 3843 [fide Guerrero-Kommritz (2001:172)]
TYPE LOCALITY. -- Zanzibar

Sepia venustoides Hoyle, 1909:266.

CURRENT SYSTEMATIC STATUS. --?Sepia pharaonis Ehrenberg, 1831 [fide Adam and Rees (1966:22)]
TYPE REPOSITORY. -- Unresolved
TYPE LOCALITY. -- Zanzibar

Sepia veranyi P. Fischer In Lagatu, 1888:115, pl 8.

CURRENT SYSTEMATIC STATUS. --?subspecies of Sepia officinalis Linne, 1758 [fide Adam and Rees (1966:151)]
TYPE REPOSITORY. -- Unresolved; Museum de Bordeaux Syntypes (2)
TYPE LOCALITY. -- Banyuls, France

Sepia vercoi Adam, 1979:190, pl 10 figs 5-6.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Reid (2000:53)]
TYPE REPOSITORY. -- WAM Holotpye 441.65 [fide Slack-Smith (1983:75)]
TYPE LOCALITY. -- West of Shark Bay, Western Australia [fide Lu (1998a:184)]

Sepia vermiculata Quoy and Gaimard, 1832:64, pl 1 figs 1-5.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:130)]
TYPE REPOSITORY. -- MNHN Holotype; specimen not extant [fide Lu et al. (1995:327)]
TYPE LOCALITY. -- Cape of Good Hope

Ascarosepion verreauxi Rochebrune, 1884b:98, pl 5 fig 2.

CURRENT SYSTEMATIC STATUS. -- Sepia mestus Gray, 1849 [fide Adam and Rees (1966:45)]
TYPE REPOSITORY. -- MNHN Syntype 1-6-175 and 1-6-176 (sepion) [fide Lu et al. (1995:327)]
TYPE LOCALITY. -- Sydney, Australia

Sepia verrucosa Lonnberg, 1896b:697.

CURRENT SYSTEMATIC STATUS. -- Sepia bertheloti D'Orbigny, 1835 [fide Adam and Rees (1966:57)]
TYPE REPOSITORY. -- ZMUU Holotype 128 [fide Wallin (1991:66)]
TYPE LOCALITY. -- Tenerife, Canary Islands

Arctosepia versuta Iredale, 1926a:194, pl 23, figs 5-6.

CURRENT SYSTEMATIC STATUS. -- Sepia limata (Iredale, 1926) [fide Reid (2000:28)]
TYPE REPOSITORY. -- AMS Holotype C133313 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Manly Beach, N.S.W., Australia

Sepia vicellius Gray, 1849:100.

CURRENT SYSTEMATIC STATUS. -- Sepia officinalis Linne, 1758 [fide Adam and Rees (1966:32)]
TYPE REPOSITORY. -- BMNH Holotype 1963120W [fide Lipinski et al. (2000:105)]
TYPE LOCALITY. -- Not designated

Sepia vietnamica Khromov, 1987a:35, fig 1.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:130)]
TYPE REPOSITORY. -- ZIN Holotype [fide Khromov et al. (1998)]
TYPE LOCALITY. -- Gulf of Tonkin (South China Sea)

Sepia (Rhombosepion) vossi Khromov, 1996:143, figs 3-4.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Norman and Lu (In press)]
TYPE REPOSITORY. -- ZMMGU Holotype Y-321 [fide Ivanov and Sysoev (2000:109)]
TYPE LOCALITY. -- East coast of Vietnam

Metasepia pfefferi wanda Iredale, 1954:78, pl 5.

CURRENT SYSTEMATIC STATUS. -- Metasepia pfefferi (Hoyle, 1885) [fide Lu (1998a:186)]
TYPE REPOSITORY. -- AMS Holotype C1333314 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Shark Bay, Western Australia

Acanthosepion whitleyanum Iredale, 1926a:195, pl 23 figs 9-10.

CURRENT SYSTEMATIC STATUS. -- Valid species; Sepia whitleyana [fide Lu (1998a:185)]
TYPE REPOSITORY. -- AMS Holotype C133331 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Port Macquarie, N.S.W., Australia

Sepia winckworthi Adam, 1939c:1, pl 1.

CURRENT SYSTEMATIC STATUS. -- Sepia brevimana Steenstrup, 1875 [fide Adam and Rees (1966:5)]
TYPE REPOSITORY. -- Unresolved; BMNH Holotype 1952.11.24.67 (sepion) [fide Adam and Rees (1966:6)]; not found [fide Lipinski et al. (2000:114)]; IRSNB Syntype [fide F.G. Hochberg (pers. comm.)]
TYPE LOCALITY. -- Singapore

Arctosepia braggi xera Iredale, 1954:74, pl 5.

CURRENT SYSTEMATIC STATUS. -- Sepia braggi Verco, 1907 [fide Lu (1998a:161)]
TYPE REPOSITORY. -- AMS Holotype C133310 [fide Rudman (1983:68)]
TYPE LOCALITY. -- Stanley, Tasmania

Sepia zanzibarica Pfeffer, 1884:9, figs 11-11a.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Khromov et al. (1998:132)]
TYPE REPOSITORY. -- Unresolved; ZMH Holotype [fide Khromov et al. (1998:132)]; no longer extant [fide Guerrero-Kommritz (2001:174)]; IRSNB Type [fide F.G. Hochberg (pers. comm.)]
TYPE LOCALITY. -- Zanzibar (East Africa)

Sepia zebrina Risso, 1854:15, pls 5-6.

CURRENT SYSTEMATIC STATUS. -- Sepia officinalis Linne, 1758 [fide Adam and Rees (1966:155)]
TYPE REPOSITORY. -- Unresolved; MHNN?
TYPE LOCALITY. -- Muddy bottoms [?Nice]

Type Repositories

AMS- Australian Museum, P.O. Box A285,† 6-8 College Street, Sydney South, New South Wales 2000, Australia.† [For type catalogs see Rudman (1983); Sweeney, Roper and Hochberg (1988) for S.S. Berry taxa]

ANSP-Academy of Natural Sciences, 19th & The Parkway, Philadelphia, PA 19103, U.S.A.† [Voss (1962e); Spamer and Bogan (1992)]

BMNH-The Natural History Museum, Cromwell Road, London SW7 5BD, England (formerly British Museum (Natural History)).† [For type catalog see Lipinski et al. (2000)]

CASIZ-California Academy of Sciences, Golden Gate Park, San Francisco, CA 94118, U.S.A.† [Smith (1974); Sweeney et al. (1988) for S.S. Berry taxa]

FCRIRM- Fisheries College and Research Institute Reference Museum Tuticorin, Tamil Nadu, India.

FMHU- Fisheries Museum, Faculty of Fisheries, Hokkaido University, Nishi 5, Kita 8, Sapporo 060, Hakodate, Japan (includes Laboratory of Marine Zoology (HUMZ) collection).† [ See Taki and Igarashi (1967)]

FMT- Taiwan Provincial Museum, 2 Siangyang Road, Taipei, Taiwan.

FUMT- University of Tokyo, Department of Fisheries, University Museum-- see MSUT.

IMC- Zoological Museum Calcutta, Indian Museum, 27 Jawaharal Nehru Road, Calcutta 700013, India [See Adam (1939a) for list of collection].

IRSNB-Institut Royal des Sciences Naturelles de Belgique, Rue Vautier 29, B-1040 Brussels, Belgium (formerly MusÈe royal d'Histoire naturelle de Belgique).

LS-† Linnaean Society of London, Burlington House, Piccadilly, London W1V 0LQ, England.† [For type catalogs see Dodge (1953, 1959)]

MHNN-Museum d'Histoire Naturelle (Musee Barla), 60 bis blvd Risso, 06300 Nice, France.

MNHN-Museum National d'Histoire Naturelle, Laboratoire Biologie Invertebres Marins et Malacologie, 55, rue de Buffon, 75005 Paris 05, France.† [For type catalog see Lu et al. (1995)]

MSNG-Museo Civico di Storia Naturale di Genova, Genova, ITALY.

MSUT-University Museum, University of Tokyo, Hongo 7-3-1, Bunkyoku, Tokyo 113, Japan (includes former Tokyo Imperial University (TIU) collection and University of Tokyo Department of Fisheries (FUMT) collection).

MZUS-MusÈe Zoologique, Universite Louis Pasteur & de la Ville de Strasbourg, 29 Boulevard de la Victoire, F67000 Strasbourg, France.† [See Johnson (1977) for Ortmann types]

NMGW- National Museums and Galleries of Wales, Cathays Park, Cardiff CF1 3NP, Wales.

NMNH-National Museum of Natural History, 10th & Constitution Avenue, Smithsonian Institution, Washington, DC 20560, U.S.A. (formerly USNM).† [For type catalogs see Roper and Sweeney (1978); Sweeney et al. (1988)]

NMP- Natal Museum, Private Bag 9070, Pietermaritzburg 3200, South Africa.

NMV- Museum of Victoria, 285-321 Russell Street, Melbourne, Victoria 3000, Australia (formerly National Museum of Victoria).† [For type catalog see Lu (1983)]

NSMT- National Science Museum, 3-23-1 Hyakunin-cho, Shinjuku-ku, Tokyo 160, Japan.† [For type catalog see Kubodera and Tsuchiya (1993)]

NTM- Northern Territory Museum of Arts and Sciences, Box 4646, Darwin, 5794, Australia.

SAM- South African Museum, Government Avenue, P.O. Box 61, Cape Town 8000, South Africa.

SAMA-South Australian Museum, North Terrace, Adelaide, South Australia 5022, Australia.† [For type catalogs see Zeidler and MacPhail (1978); Zeidler (1983); Sweeney, Roper and Hochberg (1988)]

SCSIO-South China Sea Institute of Oceanology, Academia Sinica, 164 West Xingang, Guangzhou 510301, China.

SMF- Naturmuseum und Forschungsinstitut Seckenberg, Senckenberg Anlage 25, D-60325 Frankfurt, Germany.† [For type catalog see Schafer (1938)]

WAM- Western Australian Museum, Francis Street, Perth, Western Australia 6000, Australia.† [For type catalog see Slack-Smith (1983)]

ZIN- Zoological Institute, Russian Academy of Sciences, Universitetskaya Nab. 1, St. Petersburg 199164, Russia.

ZMB- Zoologisches Museum, Museum f¸r Naturkunde der Humboldt-Universitat, Invalidenstrasse 43, D-1040 Berlin, Germany.† [For type catalog see Glaubrecht and Salcedo-Vargas (2000)]

ZMH- Zoologisches Museum, Universitat Hamburg, Martin Luther King Platz 3, 2000 Hamburg 13, Germany.† [For type catalog see Guerrero-Kommritz (2001)]

ZMMGU-Zoological Museum, Moscow State University, u1. Gensena 6, Moscow, Russia.† [For type catalog Ivanov and Sysoev (2000)]

ZMUC-Kobenhavns Universitet, Zoologisk Museum, Universitetsparken 15, DK 2100 Copenhagen, Denmark.† [For type catalog see Kristensen and Knudsen (1983); See also Volsoe et al. (1962) for Steenstrup papers]

ZMUU-Zoologiska Museet, Uppsala Universitets, Villavagen 9, S-751, 22 Uppsala, Sweden.† [For type catalog see Wallin (1991)]

ZSI- Zoological Survey of India, "M" Block, New Alipore, Calcutta 700053, India.† [For type catalog see Thomson (In prep.)]

ZSM- Zoologisches Sammlung des Bayerischen Staates, 19 Menzinger Strasse 71, 8000 Munchen 19, Bayern Germany.† [collections destroyed during WWII fide Pickford (1964)].

References

Adam, W. 1937.  Résultats scientifiques des croisières du navire-ecole Belge `Mercator' Volume I. IV-Cephalopoda.  Mémoires du Musée royal d'Histoire naturelle de Belgique, (2)9:43-82, 41 figures.

Adam, W. 1939a.  The Cephalopoda in the Indian Museum, Calcutta.  Records of the Indian Museum, 41(1):61-110, 28 figures, 2 plates.

Adam, W. 1939b.  Notes sur les Céphalopodes, XI.- Sepia bandensis sp. nov.  Bulletin du Musée royal d'Histoire naturelle de Belgique, 15(18):1-7, 4 figures.

Adam, W. 1939c.  Notes sur les Céphalopodes, XIII. - Sur une nouvelle espèce de Sepia.  Bulletin de Musée royal d'Histoire naturelle de Belgique, 15(32):1-11, 1 plate.

Adam, W. 1939e.  Siboga - Expeditie. Cephalopoda Part II. II- Revision des espèces Indo-Malaises du genre Sepia Linne, 1758. III- Revision du genre Sepiella (Gray) Steenstrup, 1880.  Results des expéditiones zoologiques, botaniques, océanographiques et entreprises aux Indes Neerlandaises Orientales en 1899-1900, 55b:35-122, 17 figures, 4 plates.

Adam, W. 1941b.  Résultats scientifiques des croisières du Navire-ecole Belge "Mercator" Volume III. IV-Cephalopoda.  Mémoires du Musée royal d'Histoire naturelle de Belgique, (2)21:83-162, 20 figures, 4 plates.

Adam, W. 1941d.  Notes sur les Céphalopodes, XVIII.- Sur les espèces de Céphalopodes de la mer Rouge décrites par C.G. Ehrenberg en 1831 et sur une nouvelle espèce de Sepia (Sepia dollfusi sp. nov.).  Bulletin du Musée royal d'Histoire naturelle de Belgique, 17(62):1-14, 2 plates.

Adam, W. 1965.  Redescription de deux espècies de Sepia insuffisamment décrites par J. Steenstrup, 1875.  Videnskabelige Meddelelser fra Dansk naturhistorisk Forening, 128:169-176.

Adam, W. 1979.  The Sepiidae (Cephalopoda: Decapoda) in the collections of the Western Australian Museum.  Records of the Western Australian Museum, 7(2):111-212.

Adam, W., and W.J. Rees. 1966.  A Review of the Cephalopod Family Sepiidae.  John Murray Expedition 1933-34, Scientific Reports, 11(1):1-165, 46 plates.

Appellof, A. 1886.  Japanska Cephalopoder.  Svenska Vetenskaps-Akademiens Handlingar, 21(13):1-36, 3 plates.

Barbut, J. 1783.  The genera Vermium exemplified by various specimens of the animals contained in the orders of the Intestina et Mollusca Linnae I. 101 pages, 11 plates. London: J. Dixwell.

Barnard, K.H. 1962.  New species and records of South African marine Mollusca from Natal, Zululand, and Mocambique.  Annals of the Natal Museum, 15(19):247-254, 4 figures.

Berry, S.S. 1912b.  A catalogue of Japanese Cephalopoda.  Proceedings of the Academy of Natural Sciences of Philadelphia, 64(2):380-444, 5 plates.

Berry, S.S. 1918.  Report on the Cephalopoda obtained by the F.I.S. "Endeavour" in the Great Australian Bight and other southern Australian localities.  Biological Results of the Fishing Experiments carried on by the F.I.S. "Endeavour," 1909-14, 4(5):201-298, 67 figures, 29 plates.

Berry, S.S. 1932.  Cephalopods of the genera Sepioloidea, Sepiadarium and Idiosepius.  The Philippine Journal of Science, 47(1):39-55, 1 plate.

Blainville, H.D.d' 1823.  Mémoire sur les espèces du genre Calmar (Loligo, Lamarck).  Journal de Physique, de Chimie et d'Histoire Naturelle, 96:116-135. [This paper (March 1823) has priority over Dict. Sci. Nat., vol 27 (July 1823)]

Blainville, H.D.d' 1824.  Mollusques, Mollusca (Malacoz.).  Dictionnaire des Sciences Naturelles, 32:1-394, atlas number 2 with 118 plates. [November 1824]

Blainville, H.D.d' 1827.  Seche, Sepia (Malacoz.).  Dictionnaire des Sciences Naturelles, 48:257-293. [July 1827]

Bosc, L.A.G. 1792.  Observation sur la Sepia rugosa (O. rugosus Fer).  Actes de la Societe d'Histoire Naturelle, Paris, 1:24.

Bosc, L.A.G. 1802.  Histoire naturelle de vers contenant leur description et leurs moeurs avec figures dessinées d'apres nature. Volume 1. Paris: Chez Deterville .. de l'imprimerie de Guilleminet. 3 volumes.

Bowdich, T.E. 1822.  Elements of conchology, including the fossil genera and the animals. Part I. Univalves. 73 pages, 19 plates. Paris: J. Smith.

Chun, C. 1915.  Die Cephalopoden. Myopsida, Octopoda.  Wissebschaftliche ergebnisse der deutschen tiefsee expedition auf dem dampfer "Valdivia" 1898-1899, 18(2):405-552, 414 figures, 34 plates.

Clarke, M.R. 1966.  A review of the systematics and ecology of oceanic squids.  Advances in Marine Biology, 4:91-300.

Cotton, B.C. 1929.  Contributions to the fauna of Rottnest Island. No. 4. Western Australian Sepiidae.  Journal of the Royal Society of Western Australia, 15:87-94, 1 figure, 3 plates.

Cotton, B.C. 1931.  Cuttlebones from Robe, with description of a new species.  South Australian Naturalist, 12(3):39-41, 9 figures.

Cotton, B.C. 1932.  Notes on Australian Mollusca with descriptions of new genera and new species.  Records of the South Australian Museum, 4(4):537-547, 9 figures.

Dewhurst, H.W. 1834.  The natural history of the Order Cetacea, and the oceanic inhabitants of the Arctic regions. 294 pages. London: Dewhurst.

Dodge, H. 1953.  A historical review of the mollusks of Linnaeus. Part 2. The Class Cephalopoda and the genera Conus and Cypraea of the Class Gastropoda.  Bulletin of the American Museum of Natural History, 103(1):1-134.

Dodge, H. 1959.  Evidential factors in the identification of the Linnaean molluscs.  Journal of the Linnaean Society of London, Zoology, 44(296):170-179.

D'Orbigny, A. 1826.  Tableau methodique de la Classe de Céphalopodes [Introduction, pages 96-120 by A.E. Ferussac].  Annales des Sciences Naturelles, Paris, (series 1)7:95-169, 245-314, 8 plates.

D'Orbigny, A. 1841-1853.  Natural History part 2, Mollusques volume 1 [1841]-1853. In R. de la Sagra, 1838-1857, Histoire physique, politique et naturelle de l'Ile de Cuba. 12 volumes and atlases. Paris.

D'Orbigny, A. 1845-1847.  Mollusques vivants et fossiles ou description de toutes les espèces de Coquilles et de Mollusques. 605 pages, atlas of 36 plates. Paris.

Ehrenberg, C.G. 1831.  Animalia invertebrata exclusis insects. Volume 4, unpaginated [126 pages].  In C.G. Ehrenberg, 1828-1845. Symbolae Physicae, seu Icones et descriptiones Corporum Nauralium novorum aut minus cognitorum... P.C. Hemprich et C.G. Ehrenberg... Pars Zoologica. 4 volumes, unpaginated. Berolini.

Ferussac, A.E. de, and A. D'Orbigny. 1834-1848.  Histoire Naturelle Générale et Particuliere Céphalopodes Acétabulifères Vivants et Fossiles. 96 pages + lvi pages + 361 pages, Atlas with 144 plates. Paris.  [See Winkworth (1942) for dates of publication]

Filippova, Y.A., and D.N. Khromov. 1991.  New data on the cuttlefish fauna (Sepiidae, Cephalopoda) from western Indian Ocean.  Zoologichesky Zhurnal, 70(8):63-70, 2 plates.

Glaubrecht, M. and M.A. Salcedo-Vargas. 2000.  Annotated type catalogue of the Cephalopoda (Mollusca) in the Museum für Naturkunde, Humboldt University of Berlin.  Mitteilungen aus dem Museum für Naturkunde Berlin, Zoologischen, 76(2):269-282.

Gmelin, J.F. 1791. Tomus I (VI), Vermes [pp. 3021-3910]. Caroli a Linne, Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima tertia, aucta, reformata. Georg. Emanuel. Beer, Lipsiae. [for publication dates see Hopkins (1908)]

Goodrich, E.S. 1896.  Report on a collection of Cephalopoda from the Calcutta Museum.  Transactions of the Linnaean Society of London, (series 2, Zoology) 7(1):1-24, 5 plates.

Gray, J.E. 1849.  Catalogue of the Mollusca in the British Museum. Part I. Cephalopoda Antepedia. 164 pages. London.

Grimpe, G. 1922.  Systematische übersicht der europäischen Cephalopoden.  Sitzungsberichte der Naturforschenden Gesellschaft zu Leipzig, 45:36-52.

Guerrero-Kommritz, J.G. 2001.  Catalogue of the cephalopod type specimens in the Zoological Museum Hamburg.  Mitteilungen. Hamburgisches Zoologische Museum and Institut, 98:171-174.

Homenko, L.P. and D.N. Khromov. 1984.  A new species of the genus Sepia (Cephalopoda, Sepiidae) from the Arabian Sea.  Zoologichesky Zhurnal, 63(8):1150-1157, 9 figures.

Hopkins, J. 1908.  Dates of publication of the separate parts of Gmelin’s edition (13th) of the ‘Systema Naturae’ of Linnaeus.  Proceedings of the Zoological Society of London, 1907:1035-1037.

Hoyle, W.E. 1885c.  Diagnosis of new species of Cephalopoda collected during the cruise of H.M.S. Challenger.- II. The Decapoda.  Annals and Magazine of Natural History, (series 5)16:181-203.

Hoyle, W.E. 1904a.  Sepia burupi n.sp. fom Natal.  Journal of Conchology, 11(1):27-28, 1 plate.

Hoyle, W.E. 1909.  A catalogue of the recent Cephalopoda, second supplement, 1897-1906.  Proceedings of the Royal Physical Society of Edinburgh, 17(6):254-299.

Hoyle, W.E. 1910b.  A list of the generic names of Dibranchiate Cephalopoda with their type species.  Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 32:407-413.

Hoyle, W.E. and R. Standen. 1901.  On a new species of Sepia and other shells collected by Dr. R. Loettlitz in Somaliland.  Memoirs and Proceedings of the Manchester Literary and Philosophical Society, 45(6):1-6.

Iredale, T. 1926a.  The cuttle-fish "bones" of the Sydney Beaches.  Australian Zoologist, 4(3):186-196, 3 figures, 3 plates.

Iredale, T. 1926b.  The biology of North-West Islet, Capricorn Group. D. Marine Mollusca.  Australian Zoologist, 4(4):237-240, 1 plate.

Iredale, T. 1940.  Marine Mollusca from Lord Howe Island, Norfolk Island, Australia, and New Caledonia.  Australian Zoologist, 9(4):429-443, 3 plates.

Iredale, T. 1954.  Cuttle-fish "Bones" again.  Australian Zoologist, 12(1):63-82, 2 plates.

Ivanov, D.L. and A.V. Sysoev. 2000.  Types of Mollusca in the Zoological Museum of Moscow University. Archives of the Zoological Museum of Moscow State University, 40:1-187 (In Russian and English).

Johnson, R.I. 1977.  Arnold Edward Ortmann, a bibliography of his work on mollusks, with a catalogue of his Recent molluscan taxa.  Occasional Papers on Mollusks, 4(58):229-244.

Jothinayagam, J.T. 1987.  Cephalopoda of the Madras Coast.  Zoological Survey of India, Technical Monograph, 15:1-85, 29 figures, 3 plates.

Keferstein, W. 1866.  Kopffusser: Cephalopoda Cuv., pages 1307-1464, plates 110-136. In H.G. Bronn, Die Klassen und Ordnungen des Thier-reiches: Weichthiere (Malacozoa). 1500 pages, 136 plates. Leipzig und Heidelberg, 1862-1866.

Khromov, D.N. 1982.  A new species of the genus Sepia (Cephalopoda, Sepiidae) from the South-West Indian Ocean.  Zoologichesky Zhurnal, 61(1):137-139, 7 figures.

Khromov, D.N. 1987a.  A new species of Sepia (Cephalopoda: Sepiidae) from the northwestern South China Sea.  Asian Marine Biology, 4:35-40.

Khromov, D.N. 1987b.  System[atics] and phylogeny of the cuttlefish family Sepiidae (Cephalopoda).  Zoologichesky Zhurnal, 66(8):1164-1176.

Khromov, D.N. 1988.  Two new species of the genus Sepia (Cephalopoda, Sepiidae) from the Sokotra Island (People's Democratic Republic of Yemen) waters.  Zoologichesky Zhurnal, 67(5):785-790, 2 figures.

Khromov, D.N. 1996.  Some notes on the shelf and slope cephalopod fauna of Vietnam, and a new species of Sepia (Cephalopoda, Sepiidae) from this region.  Ruthenica, 5(2):139-145, 4 figures.

Khromov, D.N., C.C. Lu, A. Guerra, Zh. Dong, and S.v. Boletzky. 1998.  A synopsis of Sepiidae outside Australian waters (Cephalopoda: Sepioidea), pp. 77-139. In: N.A. Voss, M. Vecchione, R.B. Toll and M.J. Sweeney (Eds.) Systematics and Biogeography of Cephalopods. Smithsonian Contributions to Zoology, 586(I-II), 599 pages.

Khromov, D.N., I.V. Nikitina and K.N. Nesis. 1991.  Three new species of cuttlefish Sepia (Doratosepion) from western Indian Ocean with new data on S. (D.) mascarensis (Cephalopoda, Sepiidae).  Zoologischesky Zhurnal, 70(6):12-24, 4 plates.

Kristensen, T.K. and J. Knudsen. 1983.  A catalogue of the type specimens of Cephalopoda (Mollusca) in the Zoological Museum, University of Copenhagen.  Steenstrupia, 9(10):217-227.

Kubodera, T. 2000a.  Cephalopoda, pages 1048-1089. In: T. Okutani, editor, Marine Mollusks in Japan. Tokai University Press, Tokyo, 1173 pages.

Kubodera, T., and K. Tsuchiya (eds.) 1993.  Catalogue of Specimens of the Class Cephalopoda (Phylum Mollusca) in the National Science Museum, Tokyo.  National Science Museum, Tokyo, 1993, unpaginated.

Lafont, A. 1868.  Extrait d'une communication faite a la Société Linneenne de Bordeaux dans la Seance du Ier Juillet.  Bulletin Hebdomadaire de l'Association scientifique de France, (series 1)4(81):120-121.

Lafont, A. 1869.  Note sur une nouvelle espèce de Sepia des cotes de France.  Journal de Conchyliologie, 17:11-14.

Lafont, A. 1871.  Note pour servir a la fauna de la Gironde contenant la liste des animaux marins dont la presence a été constatee a Arcochon pendant les années 1869-1870.  Actes de la Societe Linneenne Bordeaux, 28:237-279, 5 plates.

Lagatu, H. 1888.  Carateres distinctifs de l'espèce et du sexe dans les coquilles types de quatre Sepia.  Actes de la Societe Linneenne de Bordeaux, 42:105-120, 4 plates.

Lamarck, J.B. 1798.  Extrait d'un Mémoire sur le genre de la seche, du Calmar et Poulpe, vulgairement nommmes, Polypes de Mer.  Bulletin des Sciences, par la Societe Philomatique, 2(5):129-131.

Lesueur, C.A. 1821.  Description of several new species of cuttle-fish.  Journal of the Academy of Natural Sciences of Philadelphia, 2(1):86-101, 7 plates.

Li, F. and Q. Chen. 1989.  A new species of Sepia (Cephalopoda: Sepiidae) in the waters of the Nansha Islands of China.  Tropic Oceanology, 8(2):6-12, 4 figures.

Linne, C. 1758.  Systema naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis. 824 pages. Holmiae.

Lipinski, M.R., F.A. Naggs, and M.A. Roeleveld. 2000.  Catalogue of types of Recent cephalopods in the collection of the Natural History Museum, London.  Annales Zoologici (Warszawa), 50(1):101-120.

Lönnberg, E. 1896b.  Two cephalopods from Teneriffe collected by A. Tullgren.  Ofversigt af Kongl. Vetenskaps-Akademiens Forhandlingar, 53(10):697-706.

Lu, C.C. 1983.  The Recent Cephalopoda in the collection of the National Museum of Victoria.  Memoirs of the National Museum of Victoria, 44:65.

Lu, C.C. 1998a.  A Synopsis of Sepiidae in Australian Waters (Sepioidea: Cephalopoda), pp. 159-190. In: N.A. Voss, M. Vecchione, R.B. Toll and M.J. Sweeney (Eds.) Systematics and Biogeography of Cephalopods. Smithsonian Contributions to Zoology, 586(I-II), 599 pages.

Lu, C.C. and R. Boucher-Rodoni. 2001.  Cephalopods from the waters around Wallis and Futuna Islands in the central South Pacific.  In: P. Bouchet and B.A. Marshall (eds.), Tropical Deep-Sea Benthos volume 22. Memoires du Museum National d’Histoire Naturelle, 185:369-399, 37 figures.

Lu, C.C., R. Boucher-Rodoni and A. Tillier. 1995.  Catalogue of types of Recent Cephalopoda in the Museum national d'Histoire Naturelle (France).  Bulletin du Museum national d'Histoire naturelle, Paris, (series 4)17(3-4):307-343.

Lu, C.C. and A. Reid. 1997.  Two new cuttlefishes (Cephalopoda: Sepiidae) from the North West Shelf, and a redescription of Sepia sulcata Hoyle, 1885.  Records of the Western Australian Museum, 18:277-310, 25 figures.

Lu, C.C. and R.W. Tait. 1983.  Taxonomic studies on Sepioteuthis Blainville (Cephalopoda: Loliginidae) from the Australian region.  Proceedings of the Royal Society of Victoria, 4:181-204, 10 figures.

Mangold, K. 1998.  The Octopodinae from the eastern Atlantic Ocean and the Mediterranean Sea, pp. 521-528. In: N.A. Voss, M. Vecchione, R.B. Toll and M.J. Sweeney (Eds.) Systematics and Biogeography of Cephalopods. Smithsonian Contributions to Zoology, 586(I-II), 599 pages.

Massy, A.L. 1916.  The Cephalopoda of the Indian Museum.  Records of the Indian Museum, 12(part 5)(16):185-247, 2 plates.

Massy, A.L. 1925.  On the Cephalopoda of the Natal Museum.  Annals of the Natal Museum, 5(2):201-229, 5 plates.

Massy, A.L. 1927.  The Cephalopoda of the South African Museum.  Annals of the South African Museum, 25(4):151-167, 2 plates.

Meyer, W.T. 1909.  Cephalopoda.  Die Fauna Sudwest-Australiens, 2(19):329-335, 11 figures.

Molina, G.I. 1782.  Saggio sulla storia naturale del Chili. Bologna, Stamperia di S. Tommaso d’Aquino, 367 pages.

Molina, G.I. 1789.  Essai sur l'historie naturelle du Chili. Paris, Chez Nee de la Rochelle, 351 pages.

Naef, A. 1923.  Die Cephalopoden.  Fauna e Flora de Golfo di Napoli, Monograph 35, 1(1)(part 2):149-863, figures 63-473.

Nateewathana, A. 1996.  The Sepiidae (Cephalopoda) of the Andaman Sea, Thailand.  Phuket Marine Biological Center Special Publication, 16:145-175.

Nateewathana, A. 1998.  A new record of cuttlefish Metasepia tullbergi (Appellof, 1886) (Sepiidae: Cephalopoda), Gulf of Thailand.  Phuket Marine Biological Center Special Publication, 18(2):323-330, 1 figure.

Neethiselvan, N. 2001.  A new species of cuttlefish Sepia ramani sp. nov. (Class: Cephalopoda) from Tuticorin Bay, southeast coast of India.  Indian Journal of Marine Science, 30:81-86, 8 figures.

Nesis, K.N. 1982.  Abridged key to the cephalopod Molluscs of the world's oceans. Moscow: Light and Food Industry Publishing House, 385 pages. (In Russian; see Nesis, 1987 for abridged English translation)

Nesis, K.N. 1987.  Cephalopods of the World; Squids, cuttlefishes, Octopuses, and Allies. T.F.H. Publications, Neptune City, NJ, USA, 351 pages.

Ninni, A.P. 1884.  Catalogo dei cefalopodi dibranchiati osservati nell' Adriatico.  Atti della Societa Veneto-Trentina de Scienze Naturali in Padova, 9(1):159-174, 1 plate.

Oken, L. 1813-1826.  Okens Lehrbuch der Naturgeschichte. Volume 3, number 1, Zoologie. Leipzig.

Okutani, T. and H. Horikawa. 1987.  In: T. Okutani, M. Tagawa and H. Horikawa, Cephalopods from continental shelf and slope around Japan. 194 pages. Tokyo: Japan Fisheries Resource Conservation Association.

Okutani, T. and M. Tagawa. 1987.  In: T. Okutani, M. Tagawa and H. Horikawa, Cephalopods from continental shelf and slope around Japan. 194 pages. Tokyo: Japan Fisheries Resource Conservation Association.

Oliveira, H.P. de 1940.  Sobre dois cefalopodos da Guanabara.  Boletim do Ministerio da Agricultura, 29(10):29-35, 4 figures.

Ortmann, A. 1888.  Japanische Cephalopoden.  Zoologische Jahrbucher,(Systematik), 3:639-670, 6 plates.

Ortmann, A. 1891.  Cephalopoden von Ceylon, gesammelt von den Herren Dres. Sarasin.  Zoologische Jahrbucher, (Systematik), 5:669-678, 1 plate.

Owen, R. 1881.  Descriptions of some new and rare Cephalopoda (Part II).  Transactions of the Zoological Society of London, 11(5):131-170, 4 figures, 13 plates.

Pfeffer, G. 1884.  Die Cephalopoden des Hamburger Naturhistorischen Museums.  Abhandlungen aus dem Gebiete der Naturwissenschaften, Hamburg, 8(1):1-30, 37 figures.

Philippi, R.A. 1884.  Enumeratio molluscorum Siciliae cum viventium tum in Tellure Tertiaria fossilium quae in itinere suo observavit. Volume 2. Halis Saxonum, 303 pages, 16 plates.

Pilsbry, H.A. 1894.  Notices of new Japanese mollusks, I.  Nautilus, 7:143-144.

Quoy, J.R. and J.P. Gaimard. 1832.  Mollusques.  Voyage de decouvertes de l'Astrolabe pendant les années 1826-1827-1828-1829, Zoologie, 2(1):1-320. Atlas of 93 plates [1833].

Rafinesque, C.S. 1814.  Precis des Decouvertes et Travaux Somiologiques de Mr. C.S. Rafinesque-Schmaltz entre 1800 et 1814 ou choix raisonne de ses principales Decouvertes en Zoologie et en Botanique. 56 pages. Palermo.

Rang, M.S. 1837.  Documents pour servir a l'histoire naturelle des céphalopodes crypyobranches.  Magasin de Zoologie, 7(Cl.V.):1-77, 16 plates.

Reid, A. 1998.  A complete description of Sepia mira (Cotton, 1932) (Cephalopoda: Sepiidae) from eastern Australia.  Proceedings of the Linnanean Society of New South Wales, 119:155-171.

Reid, A. 2000.  Australian cuttlefishes (Cephalopoda, Sepiidae): the ‘doratosepion’ species complex.  Invertebrate Taxonomy, 14:1-76.

Reid, A. 2001a.  A new cuttlefish, Sepia grahami, sp. nov. (Cephalopoda: Sepiidae) from eastern Australia.  Proceedings of the Linnean Society of New South Wales, 123:159-172, 5 figures.

Reid, A. 2001b.  Sepia hedleyi Berry, 1918 (Cephalopoda: Sepiidae): a complete description and clarification of the status of S. dannevigi Berry, 1918 and S. rex (Iredale, 1926).  Records of the South Australian Museum, 34(2):79-97, 7 figures.

Reid, A. and C.C. Lu. 1998.  A new Sepiella Gary, 1849 (Cephalopoda: Sepiidae) from northern Australia, with a redescription of Sepiella weberi Adam, 1939.  The Beagle, Records of the Museums and Art Galleries of the Northern Territory, 14:71-102, 18 figures.

Risso, A. 1854.  Mollusques Céphalopodes vivants observes dans les parage méditerranéen du Comte de Nice. 81 pages, 33 plates, Nice.

Robson, G.C. 1924c.  On the Cephalopoda obtained in South African waters by Dr. J.D.F. Gilchrist in 1920-21.  Proceedings of the Zoological Society of London, 1924(2):589-686, 51 figures, 2 plates.

Robson, G.C. 1927b.  Zoological results of the Cambridge Expedition to the Seuz Canal, 1924. Report on the Mollusca (Cephalopoda).  Transactions of the Zoological Society of London, 22(3):321-329, 4 figures.

Robson, G.C. 1928c.  Céphalopodes des mers d'Indiochine.  Service Océanographique des Peches de l'Indochine, Station Maritime de Cauda, 10:1-53, 31 figures.

Robson, G.C. 1932b.  Report on the Cephalopoda in the Raffles Museum.  Bulletin Raffles Museum, 7:21-33.

Rochebrune, A.T. 1884b.  Etude monographique de la famille des Sepiadae.  Bulletin de la Societe Philomathique de Paris, (series 7)8(2):74-122, 4 plates.

Roeleveld, M.A. 1972.  A review of the Sepiidae (Cephalopoda) of southern Africa.  Annals of the South African Museum, 59(10):193-313, 20 figures, 6 plates.

Roeleveld, M.A. and W.R. Liltved. 1985.  A new species of Sepia (Cephalopoda, Sepiidae) from South Africa.  Annals of the South African Museum, 96(1):1-18, 14 figures.

Roper, C.F.E. and M.J. Sweeney. 1978.  A catalog of the type-specimens of Recent Cephalopoda in the National Museum of Natural History.  Smithsonian Contributions to Zoology, 278:1-19.

Rudman, W.B. 1983.  The cephalopod collections of the Australian Museum.  Memoirs of the National Museum of Victoria, 44:67-70.

Rüppell, E. and F.S. Leuckart. 1828.  Neue wirbellose thiere des rothen Meeres. 47 pages, 12 plates. In: Atlas zu der reise im nordlichen Afrika, von Eduard Ruppell. 1826-1828, Frankfurt am Main.

Sasaki, M. 1913a.  Decapod cephalopods found in Japan (continuation).  Dobutsu-Gaku Zasshi [Zoological Magazine Tokyo], 25(292):73-85, 12 figures, 2 plates. [In Japanese]

Sasaki, M. 1914.  Notes on the Japanese Myopsida.  Annotationes Zoologicae Japonenses, 8(5):587-629, 1 figure, 2 plates.

Sasaki, M. 1920.  Report on cephalopods collected during 1906 by the United States Bureau of Fisheries steamer "Albatross" in the northwestern Pacific.  Proceedings of the United States National Museum, 57(2310):163-203, 4 plates.

Sasaki, M. 1929.  A monograph of the dibranchiate cephalopods of the Japanese and adjacent waters.  Journal of the College of Agriculture, Hokkaido Imperial University, 20(supplement):1-357, 159 figures, 30 plates.

Schäfer, W. 1938.  Die typen de von Ruppell 1844 beschriebenen Cephalopoden.  Senckenbergiana, 20(1/2):80-94.

Schneider, J.C. 1784.  Charakteristik des ganzen geschlechts und der einzelnen arten von Blackfischen, pages 105-134. Sammlung vermischter Abhandlungen zur Aufklarung der Zoologie und der Handlungsgeschichte. 348 pages. Berlin: J.F. Unger.

Seba, A. 1734-1765.  Locupletissimi rerum naturalium thesauri accurata descriptio et iconibus artificiosissimis expressio per universam physices historiam, 1 [1734], 2 [1735], 3 [1759], and 4 [1765]. [Taxa invalid, non-binomial fide Sherborn (1902)]

Sherborn, C.D. 1902.  Index Animalium, 1758-1800. Cambridge University Press, London, 1195 pages.

Slack-Smith, S.M. 1983.  The cephalopod collection of the Western Australian Museum. Memoirs of the National Museum of Victoria, 44:71-75.

Smith, A.G. 1974.  Type and other cephalopods in the collection of the Department of Invertebrate Zoology, California Academy of Sciences.  Veliger, 16(3):319-333.

Smith, E.A. 1916.  On the shells of the South African species of the Sepiidae.  Proceedings of the Malacological Society of London, 12(1):20-26, 1 plate.

Souleyet, F.L. 1841-1852.  Mollusques, volume 2, 664 pages (1852). In: Voyage autour du monde execute pendant les années 1836 et 1837 sur la corvette La Bonite. Zoologie par MM. Eydoux et Souleyet. 2 volumes and atlas of 101 plates [atlas published 1841, but without Latin species names]. Paris.

Spamer, E.E. and A.E. Bogan. 1992.  Catalog of the types of neontological Mollusca of the Academy of Natural Sciences of Philadelphia. Part 7: Cephalopoda and Scaphopoda.  Tryonia, 24:1-13.

Steenstrup, J. 1875.  Hemisepius, en ny Slaegt af Sepia-Blacksperutternes Familie, med Bemaerkninger om Sepia-Formerne i Almindelighed.  Danske Videnskabernes Selskabs Skrifter, 5 Raekke, Naturvidenskabelig og Mathematisk, 10(7):465-482, 2 plates.

Steenstrup, J. 1881.  Sepiadarium og Idiosepius to nye Slaegter af Sepiernes Familie. Med Bemaerkninger om de to beslaegtede Former Sepiolidea D'Orb. og Spirula Lmk.  Danske Videnskabernes Selskabs Skrifter, 6 Raekke, Naturvidenskabelig og Mathematisk, 1(3):211-242, 1 plate.

Sweeney, M.J., C.F.E. Roper, and F.G. Hochberg. 1988.  Catalog of the type specimens of Recent Cephalopoda described by S. Stillman Berry.  Malacologia, 29(1):7-19.

Taki, I. and T. Igarashi. 1967.  A list of cephalopod specimens in the Fisheries Museum, Faculty of Fisheries, Hokkaido University (Sasaki collection and specimens collected by the Marine Zoological Laboratory).  Fisheries Museum, Faculty of Fisheries, Hokkaido University, Contribution, 7:1-27.

Tapparone-Canefri, C. 1877.  Contribuzioni per una fauna malacologica delle Isole Papuana. IV. Molluschi raccolti dal sig. L.M. D'Albertis nell'Isola di Sorong (Costa Nord-Ovest della Nuova Guinea) nell'anno 1872.  Annali del Museo Civico di Storia Naturale, Genova, 9:278-300.

Thiele, J. 1920.  Die Cephalopoden der deutschen Sudpolar-Expedition 1901-1903.  Deutsche Sudpolar-Expedition 1901-1903, 16(Zoology 8):433-465, 4 plates. [publication date Dec. 1920]

Thore, S. 1945.  On the Cephalopoda of Professor O. Carlgren's expedition to South Africa in 1935.  Kungliga Fysiografiska Sallskapets i Lund. Forhandlingar, 15(7):49-57, 5 figures.

Tilesius [von Tilenau], W.G. 1813.  Naturhistorische fruchte der ersten Kaiserlich - Russischen, unter dem kommando des Herrn v. Krusenstern glucklich vollbrachten erdumseeglunggesammelt. 130 pages, 5 plates, St. Petersburg.

Tillier, A., and R. Boucher-Rodoni. 1994.  Ferussac and d'Orbigny's "Histoire naturelle générale et particuliere des Céphalopodes acétabulifères": Dates of Publication of Plates and Text.  Nautilus, 107(3)[1993]:97-103. [Actual publication date 2 Feb. 1994]

Tryon, G.W. 1879.  Manual of Conchology; Structural and Systematic, with Illustrations of the Species. Volume 1, Cephalopoda, 316 pages, 112 plates. Philadelphia.

Vecchione, M., T.F. Brakoniecki, Y. Natsukari and R.T. Hanlon. 1998.  A provisional generic classification of the family Loliginidae, pp. 215-222. In: N.A. Voss, M. Vecchione, R.B. Toll and M.J. Sweeney (Eds.) Systematics and Biogeography of Cephalopods. Smithsonian Contributions to Zoology, 586(I-II), 599 pages.

Verany, J.B. 1851.  Céphalopdes de la Méditerranée.  Mollusques Méditerranéens Observes, Decrits, Figures et Chromolithographies a aprés le vivant ouvrage dedii ASM le roi Charles Albert, I:1-132, 41 plates.

Verco, J.C. 1907.  Notes on South Australian marine Mollusca, with descriptions of new species. - Part VI.  Transactions of the Royal Society of South Australia, 31:213-230, 2 plates.

Volsoe, A., J. Knudsen and W. Rees. 1962.  The cephalopod papers of Japetus Steenstrup; a translation into English. Danish Science Press, Copenhagen, 330 pages.

Voss, G.L. 1962a.  Six new species and two new subspecies of cephalopods from the Philippine Islands.  Proceedings of the Biological Society of Washington, 75:169-176.

Voss, G.L. 1962d.  South African Cephalopods.  Transactions of the Royal Society of South Africa, 36(4):245-272, 3 figures, 1 plate.

Voss, G.L. 1962e.  List of the types and species of cephalopods in the collections of the Academy of Natural Sciences of Philadelphia.  Notulae Naturae, 356:1-7.

Voss, G.L. 1963b.  Cephalopods of the Philippine Islands.  United States National Museum Bulletin, 234:1-180, 36 figures, 4 plates.

Wakiya, Y. and M. Ishikawa. 1921.  Review of myopsid cephalopods in Japan. [In Japanese]  Dobutsu-Gaku Zasshi [Zoological Magazine, Tokyo], 33:279-292, 12 figures.

Wallin, L. 1991.  Catalogue of type specimens. 2. General Zoology.  Uppsala University Zoological Museum, 80 pages.

Winkworth, R. 1936.  Marine Mollusca from South India and Ceylon. IV. A new Indian Sepia.  Proceedings of the Malacological Society of London, 22(1):16-17, 1 plate.

Winkworth, R. 1942.  Notes on the publication of Ferussac and Orbigny’s Histoire des Céphalopodes.  Proceedings of the Malacological Society of London, 25(1):34-36.

Wormuth, J.H. 1998.  Workshop deliberations on the Ommastrephidae: a brief history of their systematics and a review of the systematics, distribution and biology of the genera Martialia Rochebrune and Mabille, 1889, Todaropsis Girard, 1890, Dosidicus Steenstrup, 1857, Hyaloteuthis Gray, 1849, and Eucleoteuthis Berry, 1916, pp. 373-383. In: N.A. Voss, M. Vecchione, R.B. Toll and M.J. Sweeney (Eds.) Systematics and Biogeography of Cephalopods. Smithsonian Contributions to Zoology, 586(I-II), 599 pages.

Wülker, G. 1910.  Uber Japanische Cephalopoden. Beitrage zur kenntnis der systematik und anatomie der dibranchiaten.  Abhandlungen der mathematische-physikalische Klasse der Koeniglich Bayerischen Akademie der Wissenschaften, 3(Suppl. 1):1-77, 58 figures.

Wülker, G. 1920.  Uber Cephalopoden des Roten Meeres.  Senckenbergiana, 2(1):48-58.

Zeidler, W. 1983.  The cephalopod collection in the South Australian Museum.  Memoirs of the National Museum of Victoira, 44:77-79.

Zeidler, W. and M.K. MacPhail. 1978.  Mollusc type-specimens in the South Australian Museum, I: Cephalopoda and Scaphopoda.  Records of the South Australian Museum, 17(26):381-385.

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