Note

---

(data as of June 2002)*

Nominal Genus-Level Taxa

Asperoteuthis Nesis, 1980:59.

CURRENT SYSTEMATIC STATUS. -- Valid genus [fide Young (1992:182)]
TYPE SPECIES. -- Chiroteuthis acanthoderma Lu, 1977 by subsequent designation of Young (1992:182)

Bigelowia MacDonald and Clench, 1934:145.

CURRENT SYSTEMATIC STATUS. -- Chiroteuthis D'Orbigny, 1841 In Ferussac and D'Orbigny, 1834-1848 [fide Roper et al. (1969:11)]
TYPE SPECIES. -- Bigelowia atlanticus MacDonald and Clench, 1934 by original designation

(Chiridioteuthis) Pfeffer, 1912:554, 591.

CURRENT SYSTEMATIC STATUS. -- Chiroteuthis D'Orbigny, 1841 In Ferussac and D'Orbigny, 1834-1848 [fide Nesis (1987:241)]
TYPE SPECIES. -- Chiroteuthis pellucida Goodrich, 1896 by monotypy

Chiropsis Joubin, 1932:289.

CURRENT SYSTEMATIC STATUS. -- Chiroteuthis D'Orbigny, 1841 In Ferussac and D'Orbigny, 1834-1848 [fide Roper and Young:ToL]
TYPE SPECIES. -- Chiropsis mega Joubin, 1932 by monotypy

Chirosoma Joubin, 1912:397.

CURRENT SYSTEMATIC STATUS. -- Nomen nudum; Systematic status undetermined [fide Naef (1923:810)]
TYPE SPECIES. -- Chirosoma regnardi Joubin, 1912 by monotypy

Chiroteuthis D'Orbigny, 1841 In Ferussac and D'Orbigny, 1834-1848:xxxii.

CURRENT SYSTEMATIC STATUS. -- Valid genus [fide Nesis (1987:241)]
TYPE SPECIES. -- Loligopsis veranii Ferussac, 1834; species first mentioned, etc. [fide Hoyle (1910:408)]

(Chirothauma) Chun, 1910:240.

CURRENT SYSTEMATIC STATUS. -- Valid subgenus [fide Salcedo-Vargas (1996:22)]
TYPE SPECIES. -- Chiroteuthis imperator Chun, 1908 by monotypy

Diaphanoteuthis Tomlin, 1931:174.

CURRENT SYSTEMATIC STATUS. -- Replacement name for Leptoteuthis Verrill, 1884. Leptotheuthis Meyer, 1834, corrected to Leptoteuthis by D'Orbigny (1845:363 In 1845-1847)

Doratopsis Rochebrune, 1884:17.

CURRENT SYSTEMATIC STATUS. -- Chiroteuthis D'Orbigny, 1841 In Ferussac and D'Orbigny, 1834-1848 [fide Joubin (1900:135)]
TYPE SPECIES. -- Loligopsis vermicolaris Ruppell, 1844; subsequent designation of Hoyle (1910:408)]

Grimalditeuthis Joubin, 1898:111.

CURRENT SYSTEMATIC STATUS. -- Valid genus [fide Young (1972:75)]
TYPE SPECIES. -- Grimalditeuthis richardi Joubin, 1898 by monotypy

Leptoteuthis Verrill, 1884:140.

CURRENT SYSTEMATIC STATUS. -- Preoccupied name [fide Hoyle (1910:410)]; Chiroteuthis D'Orbigny, 1841 In Ferussac and D'Orbigny, 1834-1848 [fide Nesis (1987:241)]
TYPE SPECIES. -- Leptoteuthis diaphana Verrill, 1884 by subsequent designation of Hoyle (1910:410)]

(Planctoteuthis) Pfeffer, 1912:571, 554.

CURRENT SYSTEMATIC STATUS. -- Valid genus [fide Young, 1992:182]
TYPE SPECIES. -- Doratopsis exophthalmica Chun, 1908; species first mentioned

(Tankaia) Sasaki, 1929:308.

CURRENT SYSTEMATIC STATUS. -- Chiroteuthis D'Orbigny, 1841 In Ferussac and D'Orbigny, 1834-1848 [fide Young, 1992:182]
TYPE SPECIES. -- Chiroteuthis (Tankaia) borealis Sasaki, 1929 by original designation

Valbyteuthis Joubin, 1931:188.

CURRENT SYSTEMATIC STATUS. -- Planctoteuthis Pfeffer, 1912 [fide Young (1992:182)]
TYPE SPECIES. -- Valbyteuthis danae Joubin, 1931 by monotypy

Nominal Species-Level Taxa (As Introduced Binomial)

Chiroteuthis acanthoderma Lu, 1977:179, figs 1-14.

CURRENT SYSTEMATIC STATUS. -- Valid species; Asperoteuthis acanthoderma [fide Young (1992:182)]
TYPE REPOSITORY. -- ZMUC Holotype [fide Kristensen and Knudsen (1983:220)]
TYPE LOCALITY. -- 6°37'N, 122°02'E (Celebes Sea)

Bigelowia atlanticus MacDonald and Clench, 1934:145, figs 1-4, 6.

CURRENT SYSTEMATIC STATUS. --Chiroteuthis mega (Joubin, 1932) [fide Roper and Young: ToL]
TYPE REPOSITORY. -- MCZ Holotype 98971 [98996?]
TYPE LOCALITY. -- 37°00'N, 67°12'W (Northwest Atlantic Ocean)

Loligopsis bonplandi Verany, 1839:99, pl 1.

CURRENT SYSTEMATIC STATUS. -- Valid species; Grimalditeuthis bonplandi [fide Young (1992:174)]
TYPE REPOSITORY. -- Unresolved; MHNN? [other taxa there fide G.L. Voss (pers. comm.)]
TYPE LOCALITY. -- 29°N, 39°W (Atlantic Ocean)

Chiroteuthis (Tankaia) borealis Sasaki, 1929:309, pl 24 figs 11-14.

CURRENT SYSTEMATIC STATUS. -- Chiroteuthis sp. [fide Young (1992:182)]
TYPE REPOSITORY. -- Fisheries Institute Takashima, Hokkaido Holotype [fide Sasaki (1929:310)]
TYPE LOCALITY. -- Off Kushiro Province, Japan

Chiroteuthis calyx Young, 1972:69, pls 20, 21a-b, 22a-k, 22n-q.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Salcedo-Vargas (1996:15)]
TYPE REPOSITORY. -- SBMNH Holotype 34989 [fide Scott et al. (1990:20)]
TYPE LOCALITY. -- "Santa Catalina Basin?", eastern North Pacific Ocean

Chiroteuthis capensis Voss, 1967:76, pls 4-5.

CURRENT SYSTEMATIC STATUS. -- Chiroteuthis (Chirothauma) mega (Joubin, 1932) [fide Salcedo-Vargas (1997:97)]
TYPE REPOSITORY. -- SAM Holotype A29730
TYPE LOCALITY. -- 400 miles West of Slangkop, South Africa

Valbyteuthis danae Joubin, 1931:188, text-figs 19-32.

CURRENT SYSTEMATIC STATUS. -- Valid species; Planctoteuthis danae [fide Young (1992:169)]
TYPE REPOSITORY. -- ZMUC Holotype [fide Kristensen and Knudsen (1983:221)]
TYPE LOCALITY. -- 7°30'N, 79°19'W (Pacific Ocean)

Leptoteuthis diaphana Verrill, 1884:141, pl 32 fig 1.

CURRENT SYSTEMATIC STATUS. -- Chiroteuthis lacertosa Verrill, 1881 [fide Naef (1923:389)]
TYPE REPOSITORY. -- NMNH Holotype 38242 [fide Roper and Sweeney (1978:6)]
TYPE LOCALITY. -- 38°53'N, 69°23'W (Atlantic Ocean)

Doratopsis exophthalmica Chun, 1908:89.

CURRENT SYSTEMATIC STATUS. -- Planctoteuthis sp. [fide Young (1992:182)]
TYPE REPOSITORY. -- ZMB Lectotype Moll.-110008 [fide Glaubrecht and Salcedo-Vargas (2000:274)]
TYPE LOCALITY. -- South Indian subtropical calm zone (34°13'S, 80°30'E) [fide Chun (1910:290)]

Chiroteuthis imperator Chun, 1908:88.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Young (1992:163)]
TYPE REPOSITORY. -- ZMB Holotype Moll.-63856 [fide Glaubrecht and Salcedo-Vargas (2000:275)]
TYPE LOCALITY. -- 0°15'N, 98°08'E, Sumatra, Indonesia

Chiroteuthis joubini Voss, 1967:79.

CURRENT SYSTEMATIC STATUS. -- Valid species; Chiroteuthis (Chiroteuthis) joubini [fide Salcedo-Vargas (1996:15)]
TYPE REPOSITORY. -- ZMUC Holotype [fide Kristensen and Knudsen (1983:222)]
TYPE LOCALITY. -- 8°19'N, 44°35'W (Atlantic Ocean)

Chiroteuthis lacertosa Verrill, 1881 In 1880-1881:408, pl 56 figs 1-1f.

CURRENT SYSTEMATIC STATUS. --  Chiroteuthis veranyi [fide Roper and Young: ToL]
TYPE REPOSITORY. -- NMNH Holotype 574637 [fide Roper and Sweeney (1978:8)]
TYPE LOCALITY. -- Brown's Bank, off Nova Scotia, Atlantic Ocean (from cod stomach)

Mastigoteuthis levimana Lonnberg, 1896:605.

CURRENT SYSTEMATIC STATUS. -- Valid species; Planctoteuthis levimana [fide Young (1992:183)]
TYPE REPOSITORY. -- ZMUU Holotype 343 [fide Wallin (1991:66)]
TYPE LOCALITY. -- 43°30'N, 24°30'W (Atlantic Ocean)

Doratopsis lippula Chun, 1908:89.

CURRENT SYSTEMATIC STATUS. -- Valid species; Planctoteuthis lippula [fide Young and Roper: ToL]
TYPE REPOSITORY. -- ZMB Lectotypes Moll.-110017 [fide Glaubrecht and Salcedo-Vargas (2000:276)]
TYPE LOCALITY. -- Benguela Current (11°28'S[sic], 10°24'E) [fide Chun (1903:72)]

Cheiroteuthis macrosoma Goodrich, 1896:12, pl 3 figs 51-57.

CURRENT SYSTEMATIC STATUS. -- Chiroteuthis picteti Joubin, 1894 [fide Nesis (1987:249)]
TYPE REPOSITORY. -- ZSI Holotype M327/1[?]
TYPE LOCALITY. -- Off Kistna delta, 12°50'N, 81°30'E (Indian Ocean)

Chiropsis mega Joubin, 1932:289.

CURRENT SYSTEMATIC STATUS. -- Valid species; Chiroteuthis (Chirothauma) mega (Joubin, 1932) [fide Salcedo-Vargas (1997:95), (1996:25)]
TYPE REPOSITORY. -- ZMUC Holotype [fide Kristensen and Knudsen (1983:223)]
TYPE LOCALITY. -- 25°50'N, 76°55'W (Atlantic Ocean) [fide Joubin (1933:30)]

Valbyteuthis oligobessa Young, 1972:72, pls 23a-b, 24h-n, 24q-r.

CURRENT SYSTEMATIC STATUS. -- Valid species; Planctoteuthis oligobessa [fide Young (1992:182)]
TYPE REPOSITORY. -- SBMNH Holotype 34993 [fide Scott et al. (1990:20)]
TYPE LOCALITY. -- 32°27'N, 120°27'W, eastern North Pacific Ocean

Cheiroteuthis pellucida Goodrich, 1896:14, pl 4 figs 58-61.

CURRENT SYSTEMATIC STATUS. -- Chiroteuthis picteti Joubin, 1894 [fide Nesis (1987:249)]
TYPE REPOSITORY. -- ZSI Holotype M385/1[?]
TYPE LOCALITY. -- 16°11'15"N, 82°30'30"E (Bay of Bengal)

Loligopsis perlatus Risso, 1854:39, pl 18 figs 1-2.

CURRENT SYSTEMATIC STATUS. -- Chiroteuthis veranyi (Ferussac, 1834) [fide Adam (1952:111)]
TYPE REPOSITORY. -- Unresolved; MHNN?
TYPE LOCALITY. -- Areas of gravel [?Nice]

Onychoteuthis perlatus Risso, 1854:37, pl 17 figs 1-2.

Nomen nudum; Chiroteuthis veranyi (Ferussac, 1834) [fide Adam (1952:111)]

Chiroteuthis picteti Joubin, 1894:40, pls 1-2.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Okutani et al. (1987:151)]
TYPE REPOSITORY. -- Unresolved; MOM? Syntypes (2) [syntypes not listed by Belloc (1950:7)]
TYPE LOCALITY. -- Amboina [Indonesia]

Chiroteuthis (Planctoteuthis) planctonica Pfeffer, 1912:578, pl 46 figs 6-7.

CURRENT SYSTEMATIC STATUS. -- Undetermined
TYPE REPOSITORY. -- Unresolved; ZMH? Holotype [species not traced in Guerrero-Kommritz (2001)]
TYPE LOCALITY. -- 41.1°N, 21.1°W, Gulf Stream

Doratopsis praecox Issel, 1927:3, figs 4, 6, 7, 11-16.

CURRENT SYSTEMATIC STATUS. -- Undetermined
TYPE REPOSITORY. -- Unresolved
TYPE LOCALITY. -- Gulf of Aden

Chirosoma regnardi Joubin, 1912:396.

Nomen nudum.

Doratopsis richardi Joubin, 1912:396.

Nomen nudum.

Grimalditeuthis richardi Joubin, 1898:111.

CURRENT SYSTEMATIC STATUS. -- Grimalditeuthis bonplandi (Verany, 1839) [fide Voss (1956:151)]
TYPE REPOSITORY. -- MOM Holotype [station 750] [fide Belloc (1950:7)]
TYPE LOCALITY. -- 38°55'N, 23°39'W (E Atlantic Ocean)

Doratopsis ruppelli Rochebrune, 1884:19.

CURRENT SYSTEMATIC STATUS. -- Undetermined
TYPE REPOSITORY. -- Unresolved; MNHN? [not trace by Lu et al. (1995)]
TYPE LOCALITY. -- Straits of Messina

Doratopsis sagitta Chun, 1908:89.

CURRENT SYSTEMATIC STATUS. -- Grimalditeuthis bonplandi (Verany, 1839) [fide Young (1992:181)]
TYPE REPOSITORY. -- ZMB Lectotype Moll.-110027 [fide Glaubrecht and Salcedo-Vargas (2000:278)]
TYPE LOCALITY. -- South Indian subtropical calm zone (30°6'S, 87°50'E) [fide Glaubrecht and Salcedo-Vargas (2000:278)]

Chiroteuthis (Chirothauma) picteti somaliensis Salcedo-Vargas, 1996:22, figs 3, 5.

CURRENT SYSTEMATIC STATUS. -- Undetermined; taxon not yet reviewed
TYPE REPOSITORY. -- ZMA Holotype
TYPE LOCALITY. -- Somali Basin, Indian Ocean

Chiroteuthis (Chiroteuthis) spoeli Salcedo-Vargas, 1996:17, figs 2, 5.

CURRENT SYSTEMATIC STATUS. -- Valid species [fide Roper and Young: ToL]
TYPE REPOSITORY. -- ZMA Holotype
TYPE LOCALITY. -- Coast of Somalia, Indian Ocean

Loligopsis veranii Ferussac, 1834:355.

CURRENT SYSTEMATIC STATUS. -- Valid species; Chiroteuthis (Chiroteuthis) veranyi [fide Salcedo-Vargas (1996:15)]
TYPE REPOSITORY. -- MHNN Syntype [examined by G.L. Voss (pers. comm.)] [see also Lu et al. (1995:327)]
TYPE LOCALITY. -- Mediterranean (from dolphin stomach)

Loligopsis vermicolaris Ruppell, 1844:133 [5].

CURRENT SYSTEMATIC STATUS. -- Chiroteuthis veranyi (Ferussac, 1834) [fide Naef (1923:381)]
TYPE REPOSITORY. -- SMF Holotype; specimen not extant [fide Schafer (1938:80)]
TYPE LOCALITY. -- Messina

---

*Updated by R. Young

---

Type Repositories

MCZ- Museum of Comparative Zoology, Harvard University, Cambridge, MA 02138, U.S.A.

MHNN-Museum d'Histoire Naturelle (Musee Barla), 60 bis blvd Risso, 06300 Nice, France.

MNHN-Museum National d'Histoire Naturelle, Laboratoire Biologie Invertebres Marins et Malacologie, 55, rue de Buffon, 75005 Paris 05, France.  [For type catalog see Lu et al. (1995)]

MOM- Musée Océanographique, Avenue Saint-Martin, Monaco-Ville, MC98000, Monaco.  [For type catalog see Belloc (1950)]

NMNH-National Museum of Natural History, 10th & Constitution Avenue, Smithsonian Institution, Washington, DC 20560, U.S.A. (formerly USNM).  [For type catalogs see Roper and Sweeney (1978); Sweeney et al. (1988)]

SAM- South African Museum, Government Avenue, P.O. Box 61, Cape Town 8000, South Africa.

SBMNH-Santa Barbara Museum of Natural History, 2559 Puesta del Sol Road, Santa Barbara, CA 93105, U.S.A.  [For type catalogs see Hertz (1984); Scott et al. (1990)]

SMF- Naturmuseum und Forschungsinstitut Seckenberg, Senckenberg Anlage 25, D-60325 Frankfurt, Germany.  [For type catalog see Schafer (1938)]

ZMA- Zoologisch Museum, Universiteit van Amsterdam, P.O. Box 4766, 1009 AT Amsterdam, The Netherlands.

ZMB- Zoologisches Museum, Museum für Naturkunde der Humboldt-Universitat, Invalidenstrasse 43, D-1040 Berlin, Germany.  [For type catalog see Glaubrecht and Salcedo-Vargas (2000)]

ZMH- Zoologisches Museum, Universitat Hamburg, Martin Luther King Platz 3, 2000 Hamburg 13, Germany.  [For type catalog see Guerrero-Kommritz (2001)]

ZMUC-Kobenhavns Universitet, Zoologisk Museum, Universitetsparken 15, DK 2100 Copenhagen, Denmark.  [For type catalog see Kristensen and Knudsen (1983); See also Volsoe et al. (1962) for Steenstrup papers]

ZMUU-Zoologiska Museet, Uppsala Universitets, Villavagen 9, S-751 22 Uppsala, Sweden.  [For type catalog see Wallin (1991)]

ZSI- Zoological Survey of India, "M" Block, New Alipore, Calcutta 700053, India.  [For type catalog see Thomson (In prep.)]

References

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